Stuttgarter Beiträge zur Naturkunde Serie A (Biologie) Herausgeber: Staatliches Museum für Naturkunde, Rosenstein 1, D-70191 Stuttgart Stuttgarter Beitr. Naturk. Nr. 700 | 64S., 146 Abb. | Stuttgart, 10. TV. 2007 New species and additional records of Staphylinidae from Turkey V (Coleoptera) VOLKER ASSING Abstract 14 species of Staphylinidae from Turkey are described and illustrated: Aploderus lydicus n.sp. (Aydin, Izmir), Astenus (Eurysunius) occiduus n.sp. (Denizli, Aydin, Izmir), Medon reliquus n.sp. (Aydin, Izmir), Quedius (Raphirus) harpago n.sp. (Izmir), Cypha spathula n.sp. (Turkey: Manisa, Mersin, Osmanıye, Kahramanmaras, Antakya; Spain; Portugal), My/- laena nemorivaga n. sp. (Rize), Atheta (Datomicra) dissimulans n.sp. (Gümüshane), Drusilla lydica n.sp. (Aydin, Izmir), Pella gibbera n.sp. (Izmir), Pyroglossa pontica n.sp. (Rize), Cousya schuelkei n. sp. (Rize), Oxypoda (Oxypoda) subspectabilis n. sp. (Ordu), Dinusa smyr- nensis n.sp. (Izmir), and Pseudocalea messorphila n.sp. (Izmir). Gymnusa anatolica Korge, 1971, originally described as a subspecies of G. variegata Kiesenwetter, 1845, is regarded as a distinct species. Megalogastria Bernhauer, 1901, previously a subgenus of Aleochara Graven- horst, 1802, is elevated to genus; both the genus and its type species Aleochara cingulata Ep- pelsheim, 1889 are redescribed. A lectotype is designated for Aleochara cingulata. The male genitalia of Xantholinus osellai Bordoni (holotype), Gymnusa anatolica, and Atheta epirotica Benick are illustrated. Numerous additional records of previously described species are re- ported from Turkey, some also from Krasnodar (Russia), among them 74 first records from Turkey, one from Anatolia, one from Russia, and three from the Russian South European ter- ritory. The distributions of 14 species are mapped. Keywords: Coleoptera, Staphylinidae, taxonomy, new species, new records, lectotype de- signation, Turkey, Russia, Spain, Portugal. Zusammenfassung 14 Arten der Familie Staphylinidae aus der Türkei werden beschrieben und abgebildet: Aploderus lydicus n.sp. (Aydin, Izmir), Astenus (Eurysunius) occiduus n.sp. (Denizli, Aydin, Izmir), Medon reliquus n. sp. (Aydin, Izmir), Quedius (Raphirus) harpago n.sp. (Izmir), Cy- pha spathula n. sp. (Türkei: Manisa, Mersin, Osmaniye, Kahramanmaras, Antakya; Spanien; Portugal), Myllaena nemorivaga n.sp. (Rize), Atheta (Datomicra) dissimulans n.sp. (Gümüshane), Drusilla lydica n.sp. (Aydin, Izmir), Pella gibbera n.sp. (Izmir), Pyroglossa pontica n. sp. (Rize), Cousya schuelkei n. sp. (Rize), Oxypoda (Oxypoda) subspectabilis n. sp. (Ordu), Dinusa smyrnensis n. sp. (Izmir) und Pseudocalea messorphila n. sp. (Izmir). Gymnu- sa anatolica Korge, 1971, urspriinglich als Unterart von G. variegata Kiesenwetter, 1845 be- schrieben, wird als distinkte Art betrachtet. Megalogastria Bernhauer, 1901, bisher Untergat- tung von Aleochara Gravenhorst, 1802, wird zur Gattung erhoben und einschließlich ihrer Typusart Aleochara cingulata Eppelsheim, 1889 redeskribiert. Für Aleochara cingulata wird ein Lectotypus designiert. Die männlichen Genitalien von Xantholinus osellai Bordoni (Ho- 2 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 700 lotypus), Gymnusa anatolica und Atheta epirotica Benick werden abgebildet. Zahlreiche wei- tere beschriebene Arten werden aus der Türkei, einige auch aus Krasnodar (Russland) gemel- det, darunter 74 Erstnachweise für die Türkei, einer für Anatolien, einer für Russland und drei für das südeuropäische Russland (Russian South European territory). Die derzeit bekannte Verbreitung von 14 Arten wird anhand von Karten illustriert. Contents 199] Fats gore heres sta) re Te On ee a en Ed 2 2= Material, merbhads Sand abbreviatisns et a a ER En en er a T 3 3. Descriptions of new species and. additional records». ...2.2.2... Fide ee ec een 3 DR eterence Ss a LE Een U Le a, «Nat Oe nd en 61 1 Introduction As has been discussed in more detail earlier, the highly diverse Staphylinidae fau- na of Turkey had for a long time been neglected, but has received more attention recently (e.g. AssING 2003a, 2004a, 2004b, 2006f). Approximately 1400 species are currently known from Turkish territory; almost 200 of them were described only in the past decade (e.g. AssING 2003a, 2004a, 2004b, 2006f, 2006h, 20061, SMETANA 2004). Disregarding the Pselaphinae, the staphylinid genera with the highest diver- sity of endemic species in Turkey are Geostiba Thomson (63 species), Leptusa Kraatz (28 species and subspecies), and Sunius Curtis (25 species) (AssING 2003b, 2004e, 2006f, 2006h, 20061, in press). Unsurprisingly, numerous recently described taxa refer to these genera. However, a large number of new species have also been described in various other genera. At present, the rate of additional discoveries is still increasing rather than decreasing. The same applies to records of described - and often widespread — species which were previously unknown from Turkish ter- ritory. The present paper is based primarily on material collected during six field trips to Turkey carried out by ALEXEY SOLODOVNIKOv (currently Chicago) in June 1998, by the author in December 2005, by HEINRICH MEYBOHM (Stelle) in March 2006, by PauL WUNDERLE (Mönchengladbach) and the author in April 2006, by VOLKER BRACHAT (Geretsried) and HEINRICH MEYBOHM in April 2006, as well as by MICHAEL SCHULKE (Berlin) and the author in July/August 2006. This article is the fifth part of a series dealing with Turkish records of Staphylinidae of various genera and subfamilies. The new species and records of Leptobium Casey, Sunius, Lathro- bium Gravenhorst, Leptusa, Geostiba, and some species of Oxypoda Mannerheim have been - or will be — treated separately elsewhere (AssınG 2006a, 2006d, 2006h, 2006i, in press, in prep.). Practically all the Tachyporinae and some Oxytelinae will be studied by SCHULKE (in prep.). The present paper provides descriptions of 14 new species belonging to 14 different genera of the Oxytelinae (one species), the Paederi- nae (two species), the Staphylininae (one species), and especially of the Aleocharinae (10 species). In addition, numerous records of zoogeographic interest are compiled, among them as many as 74 first records of previously described species from Turk- ish territory. Acknowledgements My sincere thanks are due to VOLKER BRACHAT, HEINRICH MEYBOHM, MICHAEL SCHÜLKE, ALEXEY SOLODOVNIKOV, and PAUL WUNDERLE for the generous gift and loan, re- ASSING, STAPHYLINIDAE FROM TURKEY V 3 spectively, of material for the present study. JÜRGEN VOGEL (Görlitz) assisted with the iden- tification of some Atheta. I am also indepted to JOHANNES REIBNITZz (Stuttgart) for improv- ing the quality of some of the figures. 2 Material, methods, and abbreviations The material treated in the present paper is deposited in the following public institutions and private collections: cApf private collection W. Apreı, Eisenach cAss private collection V. Asstnc, Hannover cFel private collection B. FELDMANN, Münster cSch private collection M. SCHÜLkE, Berlin cSol private collection A. SOLODOVNIKOv, Chicago cWun private collection P. WUNDERLE, Mönchengladbach DEI Deutsches Entomologisches Institut, Müncheberg (L. ZERCHE) MCSNV Museo Civico di Storia Naturale Verona (L. LATELLA) NHMW Naturhistorisches Museum, Wien (H. SCHILLHAMMER) NMP Narodni Muzeum v Praze (J. Hayex) The morphological studies and drawings were carried out using a Stemi SV 11 microscope (Zeiss Germany) and a Jenalab compound microscope (Carl Zeiss Jena) with a drawing tube. For the photographs a digital camera (Nikon Coolpix 995) was used. The maps were generat- ed using the online generic mapping tool (GMT) of the Geomar website at www.aquarius. geomar.de/omc. The following morphological parameters were examined: AL length of antenna AW maximal width of abdomen EL length of elytra along suture, from apex of scutellum to posterior margin EW combined width of elytra HL head length from anterior margin of clypeus to neck HW maximal head width ML length of (median lobe of) aedeagus from base to apex of ventral process BL length of pronotum along midline PW maximal width of pronotum Tal length of metatarsus Tale length of metatibia AuB body length from mandibles to apex of abdomen The genera are arranged by subfamily; the order of subfamilies is in accordance with the checklist of Central European Staphylinidae in AssING & SCHULKE (2001). Within genera, the species are grouped according to their current subgeneric assignment or their affiliations to certain species groups. In the material sections, the Turkish provinces are arranged as follows: 1. provinces of northern Turkey from west to east; 2. provinces of western, southwestern, and southern Ana- tolia from west to east. Labels of type material are cited in their original spelling and language, except for the fol- lowing minor adaptations according to the general requirements of the journal: names of col- lectors in small capitals, scientific names of genera and species in italics, dates with the months always in Roman numbers. All the material listed in chapter 3 is from Turkey, unless stated otherwise. 3 Descriptions of new species and additional records Micropeplus fulvus Erichson, 1840 Material examined: Manisa: 1 ex., Boz Daglar, SE Turgutlu, 38°24N, 27°52E, 800 m, ceme- tery, litter of oak and shrubs, 24.X11.2005, leg. AsstnG (cAss); 1 ex., Sipil Dag Milli Park, 4 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 38°33N, 27°28E, 1080 m, 25.1V.2006, leg. BRACHAT & MEYBOHM (cAss). — Izmir: 1 ex., Boz Daglar, above Bozdag, road to ski resort, 38°21N, 28°06E, 1480m, N-slope with grass and stones, sifted, 3.IV.2006, leg. AssınG (cAss); 1 ex., Bozdag, 38°20N, 28°06E, 1300-1560 m, 21.1V.2006, leg. BRACHAT & MEYBOHM (cAss); 1 ex., Nif Dagi, 38°24N, 27°24E, 1010 m, 23.1V.2006, leg. BRACHAT & MEYBOHM (cAss). — Aydin: 4 exs., ca. 110km S Izmir, WSW Soke, Dilek Dagi, 37°39'23N, 27°08’14E, 950 m, N-slope, pine and oak litter, grass roots sift- ed, 25.X11.2005, leg. Asstnc (cAss); 1 ex., Dilek Dagi, Kanyon, 37°41N, 27°10E, 70-370 m, 16.1V.2006, leg. BRACHAT & MEYBOHM (cAss); 2 exs., Dilek Dag, S Kanyon, 37°40N, 27°11E, 670m, 17.IV.2006, leg. BRACHAT & MEYBOHM (cAss); 1 ex., ca. 20km NE Kuyucak, Bayrak Tepe, 37°58N, 28°34E, 900m, N-slope, oak litter and grass sifted, 7.TV.2006, leg. AssınG (cAss); 1 ex., NE Aydin, Imambaba Tepesi, 37°57N, 27°54E, 1460m, 20.IV.2006, leg. BRACHAT & MEYBOHM (cAss). — Mugla: 8 exs., 16km N Milas, Labranda, 37°25N, 27°43E, 670m, 21.11.2006, leg. MEYBOHM (cAss); 4 exs., Labranda, 37°25N, 27°49E, 550-660 m, 18.1V.2006, leg. BRACHAT & MEYBOHM (cAss); 1 ex., S Bafa lake, 37°28N, 27°24E, 170m, 26.11.2006, leg. MEYBOHM (cAss); 1 ex., Yatagan-Bozdogan, 37°27N, 28°18E, 825m, 19.IV.2006, leg. BRACHAT & MEYBOHM (cAss); 3 exs., Yatagan-Bozdogan, 37°38N, 28°19E, 590 m, 19.TV.2006, leg. BRACHAT & MEYBOHM (cAss); 7 exs., ca. 20 km NNE Fethiye, 36°47N, 29°11E, 970m, mixed forest, 27.III.2002, leg. AssınG (cAss); 2 exs., SE Fethiye, Baba Dag, above Ovacik, 36°33N, 29°11E, 1170 m, pine, cedar, and Quercus ilex litter, 30.111.2002, leg. AssING (cAss); 2 exs., SE Fethiye, Baba Dag, above Ovacik, 36°33N, 29°10E, 680 m, pine lit- ter, 30.111.2002, leg. AssınG (cAss); 9 exs., W Kemer, road to Ovacik, 36°36N, 30°29E, 325 m, litter of oak and other deciduous trees, 2. IV.2002, leg. AssınG (cAss). - Antalya: trex." Kalkan, Dumanlı Dagi, 36°24N, 29°26E, 1230m, forest margin, grass and moss sifted, 5.X.2002, leg. AssınG (cAss); 198 exs., ca. 20km N Kas, S Karaovabeli Gec., 36°23N, 29°43E, 830 m, paved oak and pine forest, sifted: 26.111.2002, leg. Assıng (cAss). — Mersin: 1 ex., 20 ian NNW Mut, 36°49N, 33°19E, 1320 m, pine forest with Quercus ilex, 25.X11.2000, leg. AssınG (cAss); 28 exs., road from Silifke to Gülnar, ca. 40 km W Silifke, 36°21N, 33°35E, 1015 m, oak litter sifted, 27.XII.2000, leg. AsstnG (cAss); 1 ex., road Silifke-Gülnar, 36°21N, 33°35E, 1000 m, 6.V.2004, leg. BRACHAT & MFEYBOHM (cAss); 1 ex., road to Güzeloluk, S Aydınlar, 36°42N, 34°10E, 1110 m, 5.V.2004, leg. BRACHAT & MEYBOHM (cAss); 1 ex., 23km N Silifke, 36°32N, 33°56E, 970 m, 18.1V.2005, leg. BRACHAT & MEYBOHM (cAss); 3 exs., NW Silifke, road Mut-Ermenek, 15km before Ermenek, 36°38N, 33°01E, 1030 m, 20.IV.2005, leg. BRACHAT & MEYBOHM (cAss); 1 ex., NW Tarsus, road Camlıyayla-Gözne, 37°06N, 34°37E, 570-610 m, 25.1V.2005, leg. BRACHAT & MEYBOHM (cAss). — Osmaniye: 1 ex., 15km E Os- maniye, NW Yarpuz, 37°04N, 36°26E, 920 m, stream bank, alder litter, 11.IV.2002, leg. AssınG (cAss); 1 ex., NE Kadirli, Torlar near Andirin, 37°33N, 36°26E, 1110 m, 30.IV.2005, leg. BRACHAT & MEYBOHM (cAss). — Antakya: 1 ex., 9km SE Iskenderun, 6km NE Belen, 36°32N, 36°15E, 1480 m, edge of snowfield, under stones and sifted, 4.TV.2004, leg. AssınG (cAss); 1 ex., 9km SE Iskenderun, 5km NE Belen, 36°31N, 36°15E, 1240 m, mixed oak and beech forest, sifted, 4.IV.2004, leg. Asstnc (cAss); 1 ex., Ziyaret Dagi, W Senköy, 36°01N, 36°07E, 750 m, 21.1V.2004, leg. BRACHAT & MEYBOHM (cAss); 1 ex., Ziyaret Dagi, W Sungur, 36°00N, 36°06E, 660 m, 21.1V.2004, leg. BRACHAT & MEYBOHM (cAss); 6 exs., Ziyaret Dagı, W Sungur, 35°59N, 36°05E, 710 m, 21.1V.2004, leg. BRACHAT & MEYBOHM (cAss); 1 ex., Zi- yaret Dagi, Leylekli, 35°58N, 36°03E, 510 m, 22.1V.2004, leg. BRACHAT & MEYBOHM (cAss). — Kahramanmaras: 1 ex., 20km SW Hopurlu, 520 m, 37°28’45N, 36°48’10E, 27.IV.2004, leg. BRACHAT & MEYBOHM (cAss); 1 ex., 50km W Kahramanmaras, Baskonus Yaylası, 37°34N, 36°34E, 1250 m, 5.V.2005, leg. BRACHAT & MEYBOHM (cAss). — Gaziantep: 1 ex., Kartal Dagı, W Yamacoba, 37°10N, 37°05E, 1200 m, 25.IV.2004, leg. BRACHAT & MEYBOHM (cAss). According to SMETANA (2004), this common species was previously unknown from Turkey. Micropeplus tesserula Curtis, 1828 Material examined: Mersin: 1 ex., road from Silifke to Gülnar, ca. 40km W Silifke, 36°21N, 33°35E, 1015 m, oak litter sifted, 27.XII.2000, leg. AsstNc (cAss). ASSING, STAPHYLINIDAE FROM TURKEY V 5 Micropeplus tesserula is widely distributed in the Palaearctic region, but had not been reported from Turkey (HERMAN 2001, SMETANA 2004). Proteinus meuseli Dauphin, 1995 Material examined: Izmir: 1 ex., Bozdag, 38°20N, 28°06E, 1300-1560 m, 21.IV.2006, leg. BRACHAT & MEYBOHM (cAss). The species was previously known only from Croatia, Romania, Bosnia-Herze- govina, and Greece (Assinc 2004d, DaupHIN 1995). It is here reported from Turkey for the first time. Philorinum sordidum (Stephens, 1834) Material examined: Manisa: 1 ex., Sipil Dagi Milli Parki, 38°33N, 27°28E, 1080m, 25.1V.2006, leg. BRACHAT & MEYBOHM (cAss). According to SMETANA (2004), this common species was previously unknown from Turkey. Omalium assingi Zanetti, 2002 Material examined: Manisa: 1 ex., Boz Daglar, SE Turgutlu, 38°24N, 27°52E, 800 m, ceme- tery, litter of oak and shrubs, 24.X1I.2005, leg. AssınG (cAss). This species has become known only from southern and western Anatolia, where it is apparently widespread and not uncommon, especially at higher altitudes. A dis- tribution map is provided by Assinc (2004b). Omalium cribriceps Fauvel, 1900 Material examined: Izmir: 1 ex., N Izmir, Yamanlar Dagi, 38°33N, 27°09E, 940 m, grassy patch in pine forest, under stones, sifted grass roots, 28.XII.2005, leg. AssınG (cAss); 1 ex., Nif Dagi, 38°24N, 27°24E, 1010m, 23.1V.2006, leg. BRACHAT & MEYBOHM (cAss). — Mugla: 4 exs., ca. 20km SW Mugla, N Meke, 37°13N, 28°12E, 590 m, pasture with stones at roadside, sifted and under stones, 12.IV.2006, leg. Assıng, WUNDERLE (cAss, cWun); 1 ex., ca. 25 km SW Mugla, 37°11N, 28°06E, 1130 m, N-slope, pine forest, grass roots and pine litter sifted, under stones, 12.IV.2006, leg. WUNDERLE (cWun); 2 exs., Yatagan-Bozdogan, 37°26'35N, 28°18’07E, 825 m, 19.IV.2006, leg. BRACHAT & MEYBOHM (cAss). — Ankara: 4 exs., Gölbası, 750 m, 29.X.1995, leg. Vir (cAss). In Turkey, O. cribriceps was previously known only from the Van lake area and from western Antalya province (AssınG 2004b, ZANETTI 2002). Mannerheimia brevipennis (Motschulsky, 1860) Material examined: Ankara: 8 exs., SE Ankara, N-Elma Daßı, 1600 m, moss and litter sift- ed, 31.X.1995, leg. Vrr (cAss). - Gümüshane: 4 exs., ca. 25km SW Gümüshane, Tersundagi Gec., 40°18N, 39°18E, 2070m, N-slope, spruce forest, litter and dead wood sifted, 24.VII.2006, leg. AssınG (cAss). In Turkey, this rare species is now known from Ankara, Kahramanmaras, Adıya- man, Gümüshane, and Kayseri provinces (AssING 2004b, 2006f). 6 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 Anthobium fusculum (Erichson, 1839) Material examined: Aydin: 1 ex., ca. 15 NNE km Aydin, Imambaba Tepesi, 37°57N, 27°54E, below peak 1600 m, edge of pine forest with grass, shrubs, sifted and under stones, 5.1V.2006, leg. AssınG (cAss). - Erzurum: 1 ex., 30-45km NNE Erzurum, Dumludagi, ca. 40°11N, 41°27E, 2500-2900 m, 15.V1.1998, leg. SOLODOVNIKOV (cAss). In Turkey, the species was previously known only from Mersin, Adana, and Adıyaman provinces (AssınG 2004b, 2006f). Acidota crenata (Fabricius, 1793) Material examined: Rize: 5 exs., 40km SSW Hopa, source of Caglayan D., ca. 41°06N, 41°22E, 2700-2900 m, 25.V1.1998, leg. SOLODOVNIKOV (cAss). Acidota crenata is widespread in the Palaearctic region, but was previously un- known from Turkish territory (SMETANA 2004). Aploderus lydicus n.sp. (Figs. 1-12, 50) Types Holotype 6: N37°56'47 E027°53'53 (8), Türkei, Aydin, Pasayaylasi, 1460 m, 20.1V.2006, l. BRACHAT & MEYBoHM / Holotypus 6 Aploderus lydicus sp. n. det. V. AsstNG 2006 (cAss). Paratypes: 16, 4 PP: same data as holotype (cAss); 1d: TR Prov.: Aydin (8), N Aydin, Pasayaylasi, 1460 m, 20.1V.2006, N37°56’47", E27°53'53", leg. MEYBOHM & BRACHAT (cAss); 1 2: N38°35’25 E026°29'02 (22), Türkei, Izmir, Karaburun, 550 m, 26.1V.2006, |. BRACHAT & MEYBOHM (cAss); 1 2: 28.XI1.2005, Yamanlar, 940 m, Izmir, S. ANLAs (cAss). Etymology The name (adjective) is derived from Lydia, the ancient name of the region where the species was found. Description TL: 3.5-4.5mm. Habitus as in Fig.1. Coloration: head and abdomen black; pronotum dark brown with paler lateral and posterior margins; elytra brown to dark brown; legs yellowish; antennae dark brown. Head with sexual dimorphism, larger in ¢ than in $ (Figs. 3-6); posterior sulcus absent; posterior and lateral areas with rather coarse puncturation of rather variable density, interstices on average approximately as wide as diameter of punctures; frons impunctate; integument shiny, usually with shallow microreticulation at least in pos- terior area, frons with or without transverse microstriae; eyes relatively small, ap- proximately as long as (2 and small 2) or shorter (large 3) than postocular region in dorsal view, and weakly prominent (Figs. 7-8). Antenna as in Fig. 2. Pronotum distinctly transverse, 1.35 (small 2) to 1.45 (large d) times as wide as long and 1.05 (large 3) to 1.20 (@) times as wide as head (Figs. 3-6); posterior angles completely rounded, not marked; on either side of impunctate midline with ill- defined shallow oblong impressions with coarse and dense, sometimes ill-defined puncturation; in lateral areas with less dense puncturation; integument with shallow longitudinally striate microsculpture. Elytra as wide (large d) or up to 1.1 times as wide (small 3) and at suture 1.15-1.20 times as long as pronotum (Figs. 3-6); puncturation dense and coarse, in- ASSING, STAPHYLINIDAE FROM TURKEY V / Figs. 1-12. Aploderus lydicus n. sp. — 1. Male habitus (holotype). 2. Antenna. 3-6. Forebody of three males (3-5) and female (6). 7-8. Head in lateral view ot large male (7) and small male (8). 9. Male sternite VIII. 10-12. Aedeagus in lateral and in ventral view. — Scale bars: 1 mm (1), 0.5mm (2-8), 0.2 mm (9), 0.1 mm (10-12). terstices on average narrower than diameter of punctures and without microsculp- ture. Hind wings present, length not examined. Abdomen as wide as or slightly wider than elytra, widest at segments V—-VI; punc- turation sparse and fine; tergites with distinct microreticulation; posterior margin of tergite VII broadly concave and with palisade fringe; posterior margin of tergite VIII weakly concave. 3: posterior margin of sternite VIII broadly concave (Fig.9); aedeagus as in Figs. 10-12. 8 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 Comparative notes Four Aploderus species were previously known from the Western Palaearctic re- gion, two of them widespread: the Palaearctic A. caelatus (Gravenhorst), the Euro- pean A. caesus (Erichson), A. endogaeus Assing from southwestern Anatolia (Mugla, Antalya), and the microphthalmous A. schweigeri (Smetana) from northern Anato- lia (AssING 2003a, HERMAN 2001, MAKRANCzy 2006, SMETANA 1967). In addition to the morphology of the aedeagus, the new species is distinguished from these species by the following characters: - from A. caelatus by the much smaller eyes, the more distinct and coarser punc- turation and the much shallower microsculpture of head and pronotum, as well as by the coarser puncturation of the distinctly shorter and narrower elytra; - from A. caesus by smaller average size, shorter antennae, smaller and less promi- nent eyes; the absence of a posterior sulcus on the head, the coarser, but less well de- limited puncturation of head and pronotum, the much shorter pubescence of the forebody, the absence of distinct posterior pronotal angles (in A. caesus well- marked), as well as by the shorter and narrower elytra; — from A. endogaeus by larger size, darker coloration of pronotum, elytra, and ab- domen (in A. endogaeus yellowish brown to rufous), a larger and anteriorly more distinctly widened pronotum, and by broader and longer elytra; - from A. schweigeri at once by the darker coloration, the completely different body shape (in A. schweigeri with abdomen distinctly enlarged), the distinctly larg- er eyes, more slender antennae (in A. schweigeri with distinctly transverse anten- nomeres VII-X), the more transverse pronotum, the much longer and broader ely- tra, and the presence of hind wings. For illustrations of the habitus and the genitalia of A. schweigeri and A. endogaeus (male unknown) see SMETANA (1967) and AssING (2003a). Intraspecific variation The species is subject to considerable intraspecific variation, especially in the males, affecting not only body size, but also the shape and the relative size of the head, the relative size of the eyes, the microsculpture and puncturation of the head, as well as the shape and relative size of the pronotum. For a comparison of three males of different sizes and a female see Figs. 3-8. Distribution and bionomics The species was recorded in three localities in western Turkey (Aydin and Izmir provinces) (Fig.50). Apart from the altitude (550-1460 m), no bionomic data are available. Carpelimus subtilis (Erichson, 1839) Material examined: Ankara: 2 exs., SE Ankara, N-Elma Dagi, 1300 m, hollow Salix trunk, 31.X.1995, leg. Vrr (cAss). According to SMETANA (2004), this species had not been reported from Turkey before. ASSING, STAPHYLINIDAE FROM TURKEY V 9 Oxytelus laqueatus (Marsham, 1802) Material examined: Rize: 1 ex., 30 km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1800-1900 m, 26.V1.1998, leg. SOLODOVNIKOV (cAss). Although widely distributed in the Palaearctic region and in the Americas, this species had been unknown from Turkey (HERMAN 2001). Bledius subterraneus Erichson, 1839 Material examined: Trabzon: 32 exs., ca. 50 km S Trabzon, 20km S Macka, Altindere Milli Park, 40°40N, 39°40E, 1540 m, bank of stream, 26.VII.2006, leg. Assıng, SCHULKE (cAss, cSch). According to HERMAN (2001) and SMETANA (2004), this widespread Palaearctic species had not been reported from Turkey. Stenus distortus Assing, 2006 Material examined: Gümüshane: 10 exs., ca. 50km SW Trabzon, 9-10 km S Dikkaya, ca. 40°36N, 39°29E, 2000 m, 9.V1.1998, leg. SOLODOVNIKOV (cAss, cSol). — Trabzon: 2 exs., ca. 50km S Of, S Uzungöl, 40°36N, 40°17E, 2050 m, 4.VIII.2006, leg. SCHULKE (cSch). A map illustrating the distribution of this recently described species, which has become known only from eastern and central southern Anatolia, is provided by Ass- ING (2006b). Stenus humilis Erichson, 1839 Material examined: Gümüshane: 2 exs., ca. 50km SW Trabzon, 9-10km S$ Dikkaya, ca. 40°36N, 39°29E, 2000 m, 9.V1.1998, leg. SOLODOVNIKOV (cAss, cSol). This is the first record of S. humilis from Turkey. Stenus trapezipennis Puthz, 1981 Material examined: Rize: 1 ex., 30 km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1300 m, 28.V1.1998, leg. SOLODOVNIKOV (cAss); 1 ex., 30 km SW Hopa, Caglayan river valley, ca. 41°15N, 41°13E, 500 m, 29.V1.1998, leg. SOLODOVNIKOV (cAss). This species has been recorded only from northeastern Anatolia and Georgia (HERMAN 2001). Rugilus subtilis (Erichson, 1840) Material examined: Ankara: 1 ex., SE Ankara, N-Elma Dagi, 1300 m, hollow Salix trunk, 31.X.1995, leg. Vrr (cAss). This is the first record of Rugilus subtilis trom Turkey. Rugilus lesbius Assing, 2005 Material examined: Aydin: 2 exs., Dilek Dagi, Kanyon, 37°41N, 27°10E, 70-370 m, 16.1V.2006, leg. BRACHAT & MEYBOHM (cAss). 10 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 This recently described species had been known only from the Greek island Les- bos (AssING 2005d). Astenus (Eurysunius) occiduus n.sp. (Figs. 13-21, 83) Types Holotype ¢ [with four workers of Tetramorium sp. attached to the same pin]: TR [21] - Denizli, 60 km E Mugla, S Kale, 1280 m, 37°25’39N, 28°53'18E, 11.IV.2006, V. Asstnc / Holo- typus d Astenus occiduus sp. n. det. V. AssınG 2006 (cAss). Paratypes: 1d: same data as holotype (cAss); 14, 2 2?: TR [22] - Denizli, 60km E Mugla, S Kale, 1240 m, 37°23'12N, 28°53’41E, 11.IV.2006, V. AssınG (cAss); 1 2: TR [8] - Aydin, 15km NNE Aydin, Imambaba T., below peak, 1600m, 37°57'16N, 27°53'55E, 5.1V.2006, V. Asstnc (cAss); 1 2: TR [13] - Aydin, 20 km NE Kuyucak, Bayrak Tepe, 1480 m, 38°00’09N, 28°34'53E, 7.IV.2006, P. WUNDERLE (cWun); 12: TR Prov.: Izmir (12), NO Odemis, Boz Dag, 1560 m, 21.IV.2006, N38°20'11”, E28°6'26", leg. MEYBOHM & BRACHAT (cAss). Etymology The name (Latin, adjective: western) alludes to the fact that this species is the westernmost representative of Eurysunius in Turkey. Description TL: 3.8-4.9 mm. Habitus as in Fig. 13. Coloration: forebody blackish, with the posterior 1/4-1/3 of elytra yellow, abdomen blackish with the narrow posterior mar- gins of the tergites and the apex somewhat paler, legs and antennae rufous. Head transverse, 1.25-1.30 times as wide as long; dorsal surface distinctly convex, with very dense, large, but rather shallow punctures, and only with subdued shine (Fig. 14); pubescence short, greyish, and depressed; eyes relatively small and weakly prominent, temples approximately 1.7-2.5 times as long as eyes in dorsal view (Figs. 16-17). Antenna relatively stout, antennomeres V-X moderately oblong (Fig. 15). Pronotum across anterior angles slightly (approximately 1.05 times) wider than head, 1.05-1.10 times as wide as long (width measured across anterior angles); max- imal width at anterior angles; surface without distinct impressions; posterior margin convex; lateral margins straight, each with two long setae of slightly more than half the length of lateral margin of pronotum (Fig. 14), one at anterior and one at poste- rior angle; microsculpture barely noticeable, almost absent; puncturation similar to that of head, but slightly sparser, surface somewhat more shiny than that of head; pubescence of similar length as that of head, but less fine and more conspicuous. Elytra approximately 1.1 times as wide and at suture about 0.65 times as long as pronotum; microsculpture absent; puncturation very distinct and granulose; inter- stices approximately as wide as punctures; pubescence yellowish, more distinct than that of head and pronotum; long setae present only at posterior margin near poste- rior angles, absent from lateral margins. Hind wings reduced. Abdomen about as wide as or slightly wider than elytra, widest at segments V-V]; puncturation distinct and granulose, somewhat denser on anterior than on posterior tergites; tergite VII with sparser puncturation, interstices on average twice as wide as punctures and without microsculpture; posterior margin of tergite VII with narrow rudiment of a palisade fringe. ASSING, STAPHYLINIDAE FROM TURKEY V 11 20 Figs. 13-21. Astenus occiduus n. sp. — 13. Habitus. 14. Forebody. 15. Antenna. 16-17. Head in lateral view of specimens from Denizli (16) and from Bozdag (17). 18-19. Aedeagus in lateral and in ventral view. 20-21. Apical part of aedeagus in ventral and in lateral view. - Scale bars: 1mm (13), 0.5mm (14-17), 0.2 mm (18-21). 3: sternite VII unmodified; posterior margin of sternite VIII with deep and acute incision; aedeagus as in Figs. 18-21. Comparative notes According to a recent revision of the Eurysunius species of Turkey and the Cau- casus region, three representatives were known from Turkish territory, all of them from northern and eastern Anatolia (Kastamonu, Ordu, Kayseri) (AssING 2002c). From all these species, A. occiduus is distinguished by the morphology of the aedea- gus. In addition, it is separated from them as follows: 12 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 700 - from A. bicoloratus Assıng (Ordu) by the more slender antennae (with distinct- ly oblong antennomeres V—X, the straight lateral margins of the pronotum (in A. bi- coloratus sinuate), the presence of long setae at the posterior pronotal angles, and by the absence of lateral impressions on the pronotum; - from A. paphlagonicus Assıng (Kastamonu) by less slender antennae, a trans- verse pronotum (in A. paphlagonicus about as wide as long) with sparser punctura- tion and straight lateral margins (in A. paphlagonicus weakly convex), and by the less extensive yellow coloration of the elytra (in A. paphlagonicus extending to the mid- dle of the elytra); - from A. sexsetosus Assing (Kayseri) by less slender antennae, the absence of an additional long seta between the anterior and the posterior pronotal setae, the denser pronotal puncturation, as well as by the much less extensive yellow coloration of the elytra (in A. sexsetosus extending almost to the anterior margin). For illustrations of the habitus and the genitalia of these species see AssING (2002c). Remarks Whether or not the females from Aydin and Izmir are conspecific with the types from Denizli cannot be said with certainty. The eyes of these specimens are some- what smaller (Figs. 16-17), but since no additional significant differences were found, they are here considered an expression of intra- rather than interspecific vari- ation. Distribution and bionomics The Palaearctic subgenus Eurysunius currently includes some 45 species, most of which occur in the Western Mediterranean. Only 9 species had been recorded from the Eastern Mediterranean and the Caucasus region. Astenus occiduus is the fourth representative of the subgenus to become known from Turkey and the first Eury- sunius species from Western Turkey (Denizli, Aydin, Izmir) (Fig. 83). Records of species of this subgenus are generally rare; many taxa are still represented only by their respective holotypes. Recent observations suggest that all Eurysunius species may be associated with ants of the genus Tetramorium Mayr (AssING 2002c, 2003c). This hypothesis is here supported by new evidence. At least six of the type speci- mens were found in nests of a yellowish Tetramorium sp. All the records are from grassland biotopes at altitudes of 1240-1600 m. Medon reliquus n.sp. (Figs. 22-29, 71) Types Holotype 6: TR [1] - Izmir, 1480m, Boz Daglar, above Bozdag, road to ski resort, 38°21'26N, 28°05’38E, 3.TV.2006, V. Asstnc / Holotypus d Medon reliquus sp. n. det. V. Ass- ING 2006 (cAss). Paratypes:4 dd, 6 22: same data as holotype; 2 dd, 2 2: same data, but leg. WUNDER- LE (cWun); 1d, 1 2: TR [11] - Aydin, 20km NE Kuyucak, Bayrak Tepe, 850 m, 37°57'56N, 28°33'23E, 7.1V.2006, P. WUNDERLE (cWun); 1 2: TR [12] — Aydin, 20km NE Kuyucak, Bayrak Tepe, 900 m, 37°58'06N, 28°33'33E, 7.1V.2006, P. WUNDERLE (cWun). ASSING, STAPHYLINIDAE FROM TURKEY V 13 Figs. 22-29. Medon reliquus n. sp. — 22. Habitus. 23. Forebody. 24. Antenna. 25. Head in lat- eral view. 26. Male sternite VII. 27. Male sternite VIII. 28-29. Aedeagus in lateral and in ven- tral view. — Scale bars: 1 mm (22), 0.5 mm (23-25), 0.2 mm (26-27), 0.1 mm (28-29). Etymology The name (Latin, adjective: remaining, relict) alludes to the fact that this addition to the Turkish Medon fauna was discovered only after the genus had been thoroughly revised. Description TL: 3.1-3.8 mm. Habitus as in Fig.22. Coloration: body uniformly rufous; legs and antennae pale reddish. Head approximately as long as wide to weakly oblong; puncturation dense, coarse, and well-defined; interstices on average narrower than punctures and with- out microsculpture (Fig. 23); eyes rather small (Fig. 25) and not distinctly projecting from lateral outline of head, postocular region at least 3 times as long as eyes in dor- sal view. Antenna as in Fig. 24. Pronotum approximately 1.1 times as wide as long and 1.1 times as wide as head; puncturation in lateral areas similar to that of head (Fig. 23). Elytra approximately 1.1 times as wide and at suture 0.8 times as long as prono- 14 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 tum; puncturation ill-defined, finer and denser than that of head and pronotum (Fig. 23). Hind wings reduced. Abdomen slightly wider than elytra, widest at segment VI; integument with dis- tinct microsculpture everywhere; puncturation fine and moderately dense; posterior margin of tergite VII without palisade fringe. 3: posterior margin of sternite VII almost truncate, with few modified dark setae in the middle, without palisade setae (Fig. 26); sternite VIII with relatively small pos- terior excision (Fig. 27); aedeagus as in Figs. 28-29. Comparative notes and phylogenetics The Medon species of the Western Palaearctic region were comprehensively re- vised only recently (AssING 2004c, 2006e). Medon reliquus combines an aedeagal morphology that is similar to that of some species of the M. apicalis group (M. ma- ronitus (Saulcy), M. beydaghensis Fagel) with a chaetotaxy of the male sternite VII that resembles that of species of the M. petrochilosi group (e.g. M. cerruti Coittait, M. impar Assing). This suggests that the M. apicalis group as defined earlier (AssING 2004c) may be paraphyletic and that the M. petrochilosi group should be included in the M. apicalis group. From all the species of the M. apicalis group (including those of the former M. petrochilosi group), M. reliquus is readily distinguished by the much coarser and much more well-defined puncturation of the head and pronotum (somewhat resem- bling the condition in M. brunneus (Erichson)), by the chaetotaxy of the male stern- ite VII, as well as by the shape of the aedeagus. From M. maronitus (widespread in the Eastern Mediterranean) and M. beydaghensis (southwestern Anatolia: W-Antalya, Isparta), the new species is addi- tionally separated by the uniformly reddish coloration, the absence of microsculp- ture on the forebody, the stouter modified setae at the posterior margin of the male sternite VII forming distinct patterns, as well as by the slightly different shape of the aedeagus. The geographically closest representatives of the species previously attributed to the M. petrochilosi group - all of them with restricted distributions — are M. caricus Fagel (southwestern and western Anatolia: Mugla, Izmir; Greece: Dhodhekänisos: Nikaria) and M. impar (Rhodos). From these species, M. reliquus is readily distin- guished by smaller body size, the absence of microsculpture on the forebody and consequently amuch more shiny appearance, the distinctly smaller eyes, the much shorter and more slender elytra, the less distinctly modified and less distinctly grouped setae at the posterior margin of the male sternite VII, as well as by the com- pletely different morphology of the aedeagus. For illustrations of the genitalia of the compared species see AssING (2004c). Distribution and bionomics The species is apparently endemic to the Boz Daglar and the Aydin Daglari in western Anatolia (Fig. 71). Most of the specimens were sifted from the roots of grass and shrubs, some also from gravel and from oak leaf litter on grassy north slopes and in oak forests at altitudes of 850-1480 m. The type locality is illustrated in fig. 11 in ASSING (in press a). ASSING, STAPHYLINIDAE FROM TURKEY V dey Pseudomedon obscurellus (Erichson, 1840) Material examined: Erzurum: 3 exs., 30-45km NNE Erzurum, Dumludagi, ca. 40°08N, 41°24E, 2200-2500 m, 14.V1.1998, leg. SOLODOVNIKOV (cAss, cSol). This species is widespread in Europe (SMETANA 2004). It is here reported from Turkey for the first time. Tetartopeus stylifer (Reitter, 1909) Material examined: Erzurum: 1 ex., 35km NW Tortum, Mescit Daglari, ca. 40°30N, 41°25E, 1700-2000 m, 17.V1.1998, leg. SOLODOVNIKOV (cAss). The species was previously known from Georgia, Ukraine, Russia, and Iraq (SMETANA 2004). It is here recorded from Turkey for the first time. The male geni- talia are illustrated by AssınG (2004b). Xantholinus graecus Kraatz, 1858 Material examined: Antalya: 1 ex., Alanya env., 36°32N, 32°14E, 30.V.2004, leg. WEIGEL (cAss); 1 ex., Güzelbag, Zeytinbükü river, 36°40N, 31°53E, 160 m, 24.V.2004, leg. WEIGEL (cApf); 1 ex., 15km NE Alanya, Dimcayı river, 36°34N, 32°13E, 130m, 22.V.2004, leg. WEIGEL (cApf); 1 ex., Kemer near Antalya, 800 m, 1.V.1992, leg. BEHNE (DEI). In Turkey, recent records of this species are known only from southern Anatolia (Mugla, Antalya, Mersin, Adana, Kahramanmaras, and Gaziantep provinces) (Ass- ING 2006f). Xantholinus audrasi Coiffait, 1956 Material examined: Ankara: 4 exs., Gölbası, 750m, 29.X.1995, leg. Vrr (cAss). - Ordu: 1 ex., 25km S Ordu, S Kabaduz, 40°49N, 37°54E, 990 m, road margin, roots of grass and herbs and moss sifted, 30.VII.2006, leg. AssınG (cAss). In Turkey, this species was previously reported from Mugla, Antalya, Burdur, and Mersin provinces (AssING 2003a, 2006f). Xantholinus osellai Bordoni, 1976 (Fig. 30) Type material examined: Holotype d: Turchia leg. Osetia / Draganaz gec., VI.19[year illegible] / Xantholinus osellai mihi det. BorDont 1974 / Holotypus (MCSNV). As is suggested by the similar highly derived internal structures of the aedeagus (presence of very long spine), X. osellai is the sister species of X. grandespinosus Ass- ing, from which it is distinguished as follows: body of smaller size; head smaller both absolutely and in relation to remainder of body (X. osellai: HW: 0.98 mm; HW/PW: 1.16; X. grandespinosus: HW: 1.15mm); eyes smaller, barely larger than antenno- mere I in cross-section (at widest point) (in X. grandespinosus distinctly larger); an- tennae shorter and with more transverse antennomeres, antennomeres VI-VII ap- proximately 3 times as wide as long (in X. grandespinosus about twice as wide as long); elytra without distinct microsculpture; aedeagus larger [despite smaller body size!] and with distinctly longer spine (length of spine: 1.04mm; in X. grandespinosus 0.53 mm) (Fig. 30). 16 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 Xantholinus grandespinosus Assing, 2006 Material examined: Ordu: 1 ex., 25 km S Ordu, S Kabaduz, 40°49N, 37°54E, 990 m, road margin, roots of grass and herbs and moss sifted, 30. VII.2006, leg. AssınG (cAss). Previously, only the holotype (Amasya province) of this recently described species had become known (AssING 2006f). Gabrius anatolicus Smetana, 1953 Type material examined: Holotype ¢: Yeniköy, Toros, 30.VIII.47, Anat. Exp. N. Mus. CSR. / Holotype / Gabrius Steph. anatolicus m., det. SMETANA 1952 / Mus. Nat. Pragae Inv. 18768 (NMP). Additional material examined: 4 exs., Izmir, Boz Daglar, above Bozdag, road to ski resort, 38°21N, 28°07E, 1500 m, N-slope, Alnus and Salix litter sifted, 3.1V.2006, leg. AssınG (cAss). Previously, only the type material of this species from southern Anatolia had be- come known; the above specimens represent the first record from western Anatolia. Philonthus alpinus Eppelsheim, 1875 Material examined: Rize: 3 exs., ca. 30km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1800-1900 m, 26.V1.1998, leg. SOLODOVNIKOV (cAss). — Artvin: 10 exs., ca. 40 km SW Artvin, source of Barhal river, ca. 40°05N, 41°30E, 2400m, 23.V1.1998, leg. SOLODOVNIKOV (cAss, cSol). The species is here reported from Turkey for the first time. Philonthus coprophilus Jarrige, 1949 Material examined: Erzurum: 4 exs., 30-45 km NNE Erzurum, Dumludagi, ca. 40°08N, 41°24E, 2200-2500 m, 14.V1.1998, leg. SOLODOVNIKOV (cAss, cSol). The species was previously unknown from Turkey (HERMAN 2001). Philonthus svanetiensis Coiffaıt, 1974 Material examined: Rize: 9 exs., 40km SSW Hopa, source of Caglayan D., ca. 41°06N, 41°22E, 2700-2900 m, 25.V1.1998, leg. SOLODOVNIKOV (cAss, cSol); 3 exs., ca. 30km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1800-1900 m, 26.V1.1998, leg. SOLODOVNIKOV (cAss); 1 ex., 60km SSE Rize, Ovitdagi Gec., 40°38N, 40°45E, 2510 m, N- slope, under stones, 25.VII.2006, leg. AssınG (cAss); 1 ex., Ovitdagi Gec., 40°37N, 40°47E, 2710 m, N-slope, under stones, 25. V1I.2006, leg. Asstnc (cAss). — Artvin: 1 ex., ca. 40 km SW Artvin, Barhal river valley, ca. 40°57N, 41°29E, 1800 m, 23.V1.1998, leg. SOLODOVNIKOV (cAss); 17 exs., ca. 40km SW Artvin, source of Barhal river, 41°05N, 31°30E, 2400-2800 m, 23.-24.V1.1998, leg. SOLODOVNIKOV (cAss, cSol). — Erzurum: 25 exs., 30-45 km NNE Erzu- rum, Dumludagi, ca. 40°11N, 41°27E, 2500-2900 m, 15.V1.1998, leg. SOLODOVNIKOV (cAss); 1 ex, 35km NW Tortum, Mescit Daglari, ca. 40°30N, 41°25E, 2100m, 18.V1.1998, leg. SOLODOVNIKOV (cAss); 1 ex., 35-40km NW Tortum, Mescit Daglari, ca. 40°30N, 41°17E, 2600 m, 19.V1.1998, leg. SOLODOVNIKOV (cAss). According to SMETANA (2004), the species was previously unknown from Turkey. ASSING, STAPHYLINIDAE FROM TURKEY V 17 Quedius (Raphirus) harpago n.sp. (Figs. 31-40, 50) Types _ Holotype 6: TR Prov.: Izmir (12), NO Odemis, Boz Dag, 1560m, 21.IV.2006, N38°20’11”, E28°6'26", leg. MEYBoHM & BracHat / Holotypus d Quedius harpago sp. n. det. V. AsstNG 2006 (cAss). Paratype @: N38°20’10 E028°06'25 (12), Türkei, Aydin [recte: Izmir], Bozdag, 1300-1560 m, 21.IV.2006, 1. BRACHAT & MEYBOHM (cAss). i) | * ne ie Fa Ba as am, = 4 TE g s an ee | " ve | N: A } f : T r “ La “A iJ 35 36 39 Figs. 30-40. Holotypes of Xantholinus osellai (30) and Quedius harpago n.sp. (31-39). - 30. Aedeagus. 31. Habitus. 32. Forebody. 33. Antenna. 34. Abdomen. 35-36. Median lobe of aedeagus in lateral and in ventral view. 37. Paramere. 38-39. Apical part of median lobe in ven- tral and in lateral view. 40. Internal structures of aedeagus. — Scale bars: 1mm (31-32, 34), 0.5 mm (30, 33, 35-37), 0.2 mm (38-40). 18 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 Etymology The name (Latin, noun in apposition: grapnel, grappling hook) alludes to the conspicuous shape of the apex of the median lobe of the aedeagus. Description Measurements (inmm) and ratios (holotype, paratype): AL: 2.01, -; HL: 1.10, 1.13; HW: 1.14, 1.22; PW: 1.59, 1.62; PL: 1.46, 1.53; EL: 0.84, 0.82; EW: 1.62, 1.74; AW: 1.46, 1.65; TiL: 1.19, 1.25; TaL: 1.04, 1.07; ML: 1.40, -; TL: 8.9, 8.5; HL/HW: 0.96, 0.93; PW/HW: 1.39, 1.33; PW/PL: 1.08, 1.06; EL/PL: 0.57, 0.54; EW/PW: 1.02, 1.08; AW/EW: 0.91, 0.96; TiL/TaL: 1.15, 1.17. Habitus as in Fig. 31. Coloration: body blackish; tarsi, pro- and mesofemora red- dish brown; protibia reddish brown, latero-ventrally weakly blackened; mesotibia reddish brown, latero-ventrally distinctly blackened; metafemur and metatibia blackish, with the base of the metatibia narrowly rufous; antennae reddish, with the apical half more or less infuscate, apical half of antennomere II weakly to distinctly darkened. Head weakly transverse (Fig. 32); frons without punctures; integument with very fine microsculpture composed of isodiametric meshes anteriorly and of transverse striae posteriorly. Antenna as in Fig. 33, antennomere III slightly longer than II. Pronotum distinctly wider than head and weakly transverse, with fine and shal- low transverse microsculpture, also near anterior angles. Elytra approximately as wide as and at suture only little more than half the length of pronotum (see ratios EW/PW, EL/PL, and Fig.32); puncturation distinct and dense, but rather ill-defined; microsculpture very shallow. Scutellum with 10-15 punctures. Hind wings apparently reduced. Legs relatively short; metatarsomere I approximately as long as the combined length of I-III. Abdomen widest at segment IV (Fig. 34); puncturation dense and rather fine, on- ly slightly sparser on tergite VII than on anterior tergites; anterior tergites laterally without patches of very dense pubescence; microsculpture present, but shallow; posterior margin of tergite VII with palisade fringe. 3: posterior margin of sternite VIII in the middle with rather broad (broadly V-shaped) excision; aedeagus with slender paramere reaching, but not exceeding apex of median lobe; apex of median lobe with short hook (Figs. 35-40). Comparative notes The species belongs to the group of species allied to Q. nitipennis (Stephens) of the subgenus Raphirus Stephens. It is distinguished from all its consubgeners by the morphology of the aedeagus, especially the distinctive shape of the apex of the me- dian lobe. From most similar species, it is also separated by the short elytra and the dark hind legs and from Q. nitipennis additionally by much larger size, a broader body, darker coloration, distinctly longer and more robust antennae, and coarser elytral puncturation. Distribution and bionomics The type locality is situated near the highest peak of the Boz Daglar, western Ana- tolia (Fig. 50). The absence of previous records and the short wings suggest that the species may be endemic to this mountain range, which is inhabited also by other en- demic Staphylinidae, e.g. a species of Leptobium Casey, a species of Sunius Curtis, ASSING, STAPHYLINIDAE FROM TURKEY V 19 and an undescribed species of Geostiba Thomson (AssınG 2006d, 2006h). The two type specimens were collected at an altitude of 1300-1560 m, additional bionomic data are not available. Sepedophilus binotatus (Gravenhorst, 1802) Material examined: Ankara: 1 ex., Ankara city, G. Osman Pasa, decaying tree trunk, 4.X1.1995, leg. Vrr (cAss). The species had been reported from the European part of Turkey, but was un- known from Anatolia (SMETANA 2004). Cypha spathulata n.sp. (figs. 61-69 in AssınG 2004b) Cypha squamipennis: AssınG (2004b, 2006f). Types Holotype 6: TR- Mersin, road Silifke -> Gülnar, 1015 m, No. 9, 36°20'38N, 33°35’06E, Quercus litter, 27.XI1.2000, V. Asstnc / Holotypus d Cypha spathulata sp. n. det. V. AssING 2006 (cAss). Paratypes: 1d: TR - Manisa [4], 1000m, Boz Daglar, SE Turgutlu, N-slope with oak, 38°22'26N, 27°50'53E, 24.XI1I.2005, V. Asstnc; 1 2: TR [50], 1250m, W Kahramanmaras, Baskonus Yaylası, 5.V.2005, 37°33'56N, 36°33'37E, BRACHAT & MEYBOHM (cAss); 1d: TR- Mersin, 25, 1085m, Camliyayla, Quercus & Carpinus near creek, 37°10'24N, 34°36’35E, 5.V.2002, MEYBOHM (cAss); 16: TR [45] -— Osmaniye, 12.5km NE Andırın, -> Geben, 37°39'14N, 36°26'27E, 1500 m, 3.V.2005, BRACHAT & MEYBOHM (cAss); 1: Zıyaret Dagı, 19km S Antakya, SW Senköy, 36°01'48N, 36°07’19E, 913 m, oak & laurel shrubs, sifted, 2.1V.2004, leg. M. SCHULKE (cSch). — SPAIN: 2 dd, 2 22: E- Andalusia (CA), Grazalem- ma, 1220 m, 2.X.93, WUNDERLE (cAss, cWun); 3 dd: E — Andalusia (M) [sic], Cortez Fron- terra [sic], 2.X.93, WUNDERLE / Sierra Cortez de Fronterra [sic], 1200 m, Steineiche (cAss, cWun); 18, 522: E — Andalusia, Sierra Nevada, 28.1X.93, WUNDERLE / Guejar Sierra, 1200 m, Genilufer, Bachmoos, Gesiebe (cCWun). — PORTUGAL: 1 6: P - Algarve, 400 m, 10km N S. Bras, 4.V1.1992, WUNDERLE (cWun). Etymology The name (Latin, adjective derived from the noun spathula) alludes to the characteristic shape of the apex of the ventral process of the aedeagus. Description Measurements (in mm) and ratios (range, arithmetic mean; n = 10): AL: 0.44-0.60, 0.55; HW: 0.36-0.44, 0.41; PW: 0.53-0.68, 0.63; PL: 0.29-0.36, 0.34; EL: 0.30-0.41, 0.37; EW: 0.57-0.72, 0.67; AW: 0.51-0.62, 0.58; TiL: 0.26-0.36, 0.33; TaL: 0.15-0.23, 0.20; ML: 0.32-0.36, 0.35; TL: 1.3-1.9, 1.65; PW/HW: 1.41-1.62, 1.51; PW/PL: 1.75-1.91, 1.83; EL/PL: 1.04-1.13, 1.08; EW/PW: 1.02-1.12, 1.09; AW/EW: 0.83-0.89, 0.86; TiL/TaL: 1.53-1.85, 1.68. Habitus as in figs. 61-62 in AssiNG (2004b). Coloration variable: head and prono- tum brown to dark brown, the latter with broad yellowish margins; elytra light brown to brown; abdomen brown to dark brown, with the posterior halves of seg- ments III-VI and all of segments VII-IX at least slightly paler; legs, palpi, and an- tennae bright yellow to light brown. Head and pronotum with sparse and extremely fine, barely noticeable punctura- tion; microsculpture indistinct, pubescence depressed. Antenna with antennomere I large and weakly oblong; II of similar length but distinctly narrower, almost twice as 20 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 long as wide; III-V minute, much narrower than II, of subequal length, and more or less distinctly oblong; VI slightly larger and longer than V; VII similar to VI, but slightly shorter; VIII-X forming distinct club, of similar width; VIII and IX weakly oblong; XI slightly longer than the combined length of VIII and IX (figs. 64-65 in AssING 2004b). Pronotum with posterior angles indistinct, completely rounded; distinctly trans- verse (see ratios PL/PW and PW/HW). Elytra wider and at suture usually slightly longer than pronotum (see ratios EW/PW, EL/PL); puncturation as fine as that of pronotum; microsculpture distinct, composed of diagonal striae and long meshes. Hind wings present. Metatarsomere I almost as long as the combined length of the following tarsomeres or slightly short- er. Abdomen with distinct microsculpture and extremely fine sparse puncturation; posterior margin of tergite VII with palisade fringe. 3: protarsomere I moderately dilated, slightly longer than the combined length of II-III; aedeagus of distinctive shape, ventral process apically spathuliform (figs. 66-69 in AssınG 2004b). Comparative notes The species is separated from all its congeners by the distinctive morphology of the aedeagus. From the two other Cypha species recorded from Turkey, C. laevius- cula (Mannerheim) and C. tenebricosa Assing, it is easily distinguished also by the paler coloration, especially of the legs (in C. laeviuscula and C. tenebricosa dark brown to black). The external and sexual characters of C. tenebricosa are figured by AssING (2004b). For illustrations of the male genitalia and antennae of many Euro- pean species see DAUPHIN (2004) and PaLm (1936). Intraspecific variation Considerable intraspecific variation was observed particularly regarding size (a nanistic specimen is illustrated in figs. 62, 63, 65, 67, 69 in AssING 2004b), col- oration (especially legs and antennae), length and proportions of antennae, width of body, proportions of pronotum and elytra, and the size of the median lobe of the aedeagus. In the material from southwestern Europe, the apex of the median lobe of the aedeagus is slightly more strongly dilated than in the Turkish specimens, but this difference is attributed to intra- rather than interspecific variation. Comments Most of the type material listed above was previously interpreted and recorded as C. squamipennis (Fauvel) (AssiNG 2004b, 2006f). Quite recently, however, PATRICK DauPHIN (pers. comm.), who studied and dissected the male holotype of Cypha squamipennis and prepared a drawing of the aedeagus, kindly informed me that the redescription and figures given in AssING (2004b) referred to a different species, not to C. squamipennis. Distribution and bionomics The species is recorded from Turkey (Manisa, Mersin, Osmaniye, Kahraman- maras, Antakya), Spain, and Portugal, so that it can be expected to be present in most ASSING, STAPHYLINIDAE FROM TURKEY V 21 or all of the Mediterranean. The types were mostly sifted from the leaf litter of de- ciduous trees at altitudes of 400-1540 m. Oligota pumilio Kiesenwetter, 1858 Material examined: Ankara: 1 ex., SE Ankara, N-Elma Dagi, 1200m, Crataegus litter, 31.X.1995, leg. Vir (cAss); 24 exs. [1 2 macropterous, remainder with wings of reduced length], SE Ankara, N-Elma Dagi, 1300 m, decaying and hollow Salix trunks, 31.X.1995, leg. Vit (cAss). This species was only recently reported from Turkey (Antalya and Kahraman- maras provinces) for the first time (AssING 2006f). Only one of the examined speci- mens is macropterous; the remainder has wings of reduced length. Oligota inversa Assing, 2002 Material examined: Izmir: 1 ex., N Izmir, Yamanlar Dagi, 38°33N, 27°10E, 680m, 28.X1I.2005, leg. Assıng (cAss). — Ankara: 1 ex., SE Ankara, N-Elma Dagi, 1200 m, Cratae- gus litter, 31.X.1995, leg. Vrr (cAss). Oligota inversa was previously known only from Greece (AssING 2002b). Oligota anatolica Assing, 2003 Material examined: Izmir: 7 exs., 20 km E Izmir, Nif Dag, 38°24N, 27°24E, 920 m, plateau, sifted grass roots between stones under shrubs, 26.XT1.2005, leg. AssınG (cAss). This is the westernmost record of O. anatolica, which was previously known from Denizli, Antalya, Mersin, Kahramanmaras, and Adıyaman provinces, as well as from Israel (AssING 2003a, 2006f; Kapp 2004). Myllaena infuscata Kraatz, 1853 Material examined: Mugla: 69 exs., Gölgeli Daglari, 20km NE Köycegiz, above Agla, 37°03N, 28°49E, 1690 m, grass, thistles, and pine litter sifted, 6.X.2002, leg. AssınG (cAss). - Osmaniye: 3 exs., Osmaniye, NE Kadirli, 10 km N Andirin-Cokak, 37°39N, 36°21E, 1150 m, 1.-2.V.2005, leg. BRACHAT & MEYBOHM (cAss). — Adıyaman: 1 ex., ca. 50 km NE Adiyaman, ca. 5km N Sincik, 38°03N, 38°37E, 1320 m, under stones near stream, 23.111.2005, leg. Assinc (cAss). Myllaena infuscata is widespread in the Western Palaearctic region, but had not been reported from Turkey (SMETANA 2004). Myllaena nemorivaga n.sp. (Figs. 41-48, 50) Types Holotype d: TR [23] - Rize, 25km SSE Rize, 4km E Ikizdere, 750m, 40°47’14N, 40°35’31E, 31.VII.2006, V. AssınG / Holotypus 3 Myllaena nemorivaga sp. n. det. V. AssING 2006 (cAss). Paratypes: 16 dd, 13 2¢: same data as holotype (cAss); 60 exs.: same data, but leg. M. SCHULKE (cSch); 8 dd, 6 22: same data, but “[23a] ... 1.VIII.2006, V. Asstnc” (cAss); 57 exs.: same data, but leg. M. SCHULKE (cSch); 18,2 29: TR [24] - Rize, 25 km SSE Rize, 7km E Ik- izdere, 1030 m, 40°47’01N, 40°38’18E, 31.VII.2006, V. Asstnc (cAss); 1 ex.: same data, but leg. 22 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 44 Figs. 41-49. Myllaena nemorivaga n. sp. (41-48) and Gymnusa anatolica (49). — 41. Habitus. 42. Forebody. 43. Antenna. 44. Head in lateral view. 45-47. Median lobe of aedeagus in later- al and in ventral view. 48. Spermatheca. 49. Median lobe of aedeagus in lateral view. — Scale bars: 1 mm (41), 0.5 mm (42-44, 49), 0.1 mm (45-47), 0.05 mm (48). M. SCHULKE (cSch); 8 exs.: TR [29] — Rize, 25 km SE Rize, 900m, stream bank, Rhododen- dron, 40°53'32N, 40°46’03E, 2.VIII.2006, M. SCHULKE (cSch). Etymology The name (Latin, adjective: wandering in the forest) alludes to the fact that, in contrast to most of its congeners, the species is apparently an inhabitant of the forest floor. Description TL: 2.2-2.8mm. Habitus as in Fig.41. Coloration: forebody rufous to reddish brown, with the head sometimes slightly darker; abdomen brown to dark brown, with segment VIII and following yellowish to yellowish brown; legs and antennae dark yellowish to yellowish brown. ASSING, STAPHYLINIDAE FROM TURKEY V 23 Head with eyes of reduced size, composed only of few (< 10) ommatidia (Fig. 44), not distinctly projecting from lateral outline of head, and only approximately !/4 the length of postocular region in dorsal view. Antenna of moderate length; anten- nomeres IV-VI moderately to weakly oblong; VII-IX weakly oblong or as long as wide; X as long as broad or weakly transverse; XI shorter than the combined length of IX-X (Fig. 43). Pronotum strongly convex in cross-section, approximately 1.25 times as wide as long and 1.5 times as wide as head; maximal width in posterior half, more strongly tapering cephalad than caudad; posterior angles distinct and subrectangular; posteri- or margin near posterior angles indistinctly sinuate at most (Fig. 42). Elytra conspicuously short, approximately as wide and at suture only 0.50-0.55 times as long as pronotum; posterior margin distinctly sinuate near posterior angles (Fig. 42); puncturation dense and fine, but more distinct than the extremely fine and dense puncturation of the head and pronotum. Hind wings reduced. Metatarsomere I as long as the combined length of II-III, or slightly shorter. Abdominal tergite VII with or without narrow rudiment of a palisade fringe at the posterior margin; tergite VIII without sexual dimorphism, its posterior margin strongly convex in both sexes. 3: posterior margin of sternite VIII obtusely angled in the middle; median lobe of aedeagus of distinctive shape and with distinctive internal structures (Figs. 45-47). ?: sternite VIII broadly convex posteriorly; spermatheca as in Fig. 48, duct with approximately 8 coils. Comparative notes From other Western Palaearctic Myllaena species, M. nemorivaga is distinguished not only by the highly distinctive genitalia, but also by the following combination of external characters: reddish to reddish brown coloration, relatively small size, eyes of strongly reduced size (< 10 ommatidia), pronotum with marked posterior angles, very short elytra, and reduced hind wings. Based on the original description Fig.50. Distributions of Aploderus lydicus n. sp. (@), Myllaena nemorivaga n. sp. (©), Quedius harpago n.sp. (OD), and Atheta dissimulans n. sp. (M). 24 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 (EPPELSHEIM 1880), M. caucasica, which has also been recorded from Turkey, is sep- arated from the new species by darker coloration (pronotum and elytra dark brown to blackish brown), the longer antennae (all antennomeres oblong), the rounded posterior angles of the pronotum, and the longer elytra. Distribution and bionomics The types were found in the area to the east of Ikizdere, Rize province (Fig. 50). They were collected by sifting the leaf litter of mixed forests composed mainly of chestnut, alder, beech, and Rhododendron at altitudes of 750 and 1030m. Some of the paratypes are teneral. Gymnusa anatolica Korge, 1971; n. stat. (Fig. 49) Gymnusa variegata anatolica Korge, 1971; KoRGeE (1971: 59f.). Material examined: Gümüshane: 3 exs., ca. 30km ENE Gümüshane, Kostandagı Geg., 40°31N, 39°47E, 2340 m, N-slope, 24.VII.2006, leg. Assıng, SCHULKE (cAss, cSch). Based on a single teneral holotype female from the surroundings of Rize, KORGE (1971) tentatively described this taxon as a subspecies of G. variegata Kiesenwetter. To my knowledge, no further records had become known. A comparison of the above specimens with material of G. variegata revealed no differences in coloration; the differences pointed out in the original description are artefacts resulting from the teneral condition of the holotype. However, the other distinguishing characters (puncturation of the abdomen, shape of the posterior margin of tergite VIII) are confirmed. Moreover, the aedeagus is of different shape (Fig. 49): the median lobe is larger, the conspicuous process at the base of the ventral process is longer, more slen- der, and apically more acute, and the ventral process is longer and apically more strongly bent. Transitional conditions are unknown and there are records of G. variegata neither from the southern Balkans, nor from other regions in Turkey, nor from the Caucasus region. Thus, both the morphological and the zoogeographic ev- idence suggests that G. anatolica represents a distinct species rather than a subspecies of G. variegata. Gyrophaena gentilis Erichson, 1839 Material examined: Ordu: 11 exs., 25km S Ordu, S Kabaduz, 40°49N, 37°54E, 990 m, mixed forest with alder, spruce, bramble, ivy, 30.VII.2006, leg. AssING, SCHULKE (cAss, cSch). — Giresun: 2 exs., ca. 30km S Giresun, 40°35N, 38°27E, 1350 m, spruce forest with Rhodo- dendron, 29.VI1.2006, leg. Assıng, SCHULKE (cAss, cSch). — Trabzon: 10 exs., ca. 50km S Trabzon, 20km S Macka, Altindere Milli Park, 40°40N, 39°40E, 1560 m, spruce forest with Rhododendron, sifted, 26.V1I.2006, leg. AssING, SCHULKE (cAss, cSch); 1 ex., Altindere Milli Park, 40°41N, 39°39E, 1650 m, spruce forest with Rhododendron, sifted, 26.VII.2006, leg. SCHULKE (cSch). — Rize: 5 exs., ca. 30 km S Ardesen, 40°56N, 40°58E, 750 m, moist forest with boxwood and Rhododendron, sifted, 3.VIII.2006, leg. SCHULKE (cSch). According to SMETANA (2004), this widespread Palaearctic species was unknown from Turkey. Rhopalocerina clavigera (Scriba, 1859) Material examined: Rize: 2 exs., ca. 30km S Ardesen, 40°56N, 40°58E, 750 m, moist forest with boxwood and Rhododendron, sifted, 3.VIII.2006, leg. SCHULKE (cAss, cSch); 1 ex., ca. ASSING, STAPHYLINIDAE FROM TURKEY V 25 50km S Ardesen, Cat, 40°52N, 40°56E, 1240 m, alder forest, sifted, 3. VIII.2006, leg. SCHULKE (cSch). Rhopalocerina clavigera is widespread in Europe, but had not been reported from Turkey (SMETANA 2004). Bolitochara obliqua Erichson, 1837 Material examined: Sinop: 1 ex., Cangal Dagi, 7.-15.V1.1960, leg. SCHUBERT (cAss). — Tunceli: 1 ex., Ovacik, 1400 m, VI.1976, leg. SCHUBERT (NHMW). — Mersin: 1 ex., Camli- yayla, 10.V.-3.V1.1963, leg. SCHUBERT (NHMW). - Bitlis: 1 ex., Bitlis, 1700 m, VI.1971, leg. SCHUBERT (NHMW); 1 ex., Tatvan, 1900 m [without date], leg. SCHUBERT (cAss). This widespread and common species is here reported from Turkish territory for the first time. Bolitochara lauferi Bernhauer, 1908 Material examined: Izmir: 1 ex., 20km E Izmir, Nif Dagi, 38°23N, 27°22E, 1270-1400 m, Pinus litter and grass roots sifted, 26.XTI.2005, leg. AssınG (cAss). Bolitochara lauferi was previously known only from central southern Anatolia (AssING 2006f). Myrmecopora convexula Assing, 1997 Material examined: Izmir: 1 ex., N Izmir, Yamanlar Dagi, 38°33N, 27°10E, 680m, road margin, in nest of Messor sp., 28.XII.2005, leg. AssınG (cAss); 6 exs., ca. 80km NW Izmir, W Karaburun, 38°38N, 26°29E, 440m, pasture with stones on limestone, nest of Messor sp., 4.1V.2006, leg. Assıng, WUNDERLE (cAss, cWun). — Manisa: 3 exs., Boz Daglar, ca. 10 km WSW Turgutlu, Sivrice, 38°27N, 27°50E, 330m, ruderal meadow, nest of Messor sp., 4.IV.2006, leg. AssınG (cAss). This myrmecophilous species has become known only from the surroundings of Izmir and from the Greek island Lesbos (Assıng 1997b, 2005d). Cordalia obscura (Gravenhorst, 1802) Material examined: Zonguldak: 1 ex., Amaclar env., 190m, 17.-18.V1.2003, leg. SMATANA (cAss). Though generally widespread and common in the Western Palaearctic region, this species has rarely been reported from Turkey. Since previous records may be based on a confusion with other Cordalia species occurring in Turkey (C. anatolica Ass- ing, C. fortepunctata Assing, C. rose: Assing), which were described only very re- cently, the above specimen represents the first confirmed Turkish record of C. ob- scura. Cordalia fortepunctata Assing, 2006 Material examined: Ankara: 1 ex., SE Ankara, N-Elma Dagi, 1300 m, hollow Salix trunk, 31.X.1995, leg. Vir (cAss). This very recently described species was previously known from Mersin, Kahra- manmaras, Kayseri, Malatya, and Adıyaman provinces (AssING 2006f). 26 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 Falagrioma thoracica (Stephens, 1832) Material examined: Ordu: 1 ex., 25 km S Ordu, S Kabaduz, 40°49N, 37°54E, 990 m, mixed forest with alder, spruce, bramble, ivy, 30.VII.2006, leg. AssınG (cAss). SMETANA (2004) does not list the widespread Western Palaearctic F thoracica for Turkey, so that this is apparently the first record. Autalia puncticollis Sharp, 1864 Material examined: Rize: 2 exs., 30km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1300 m, in mushrooms, 28.V1.1998, leg. SOLODOVNIKOV (cAss). Autalia puncticollis is widespread in the Western Palaearctic region, except for the extreme southwest and the north (AssiNG 1997a); the record from Japan (SMETANA 2004) is very likely to be based on a misidentification. The species is here recorded from Turkey for the first time. Ischnopoda ulbrichi Lohse, 1994 Material examined: Trabzon: 2 exs., ca. 50km S Trabzon, 20km S Macka, Altindere Milli Park, 40°40N, 39°40E, 1540 m, bank of stream, 26.VII.2006, leg. AsstNG (cAss). According to a recent revision of the genus (Pasnık 2006), I. ulbrichi was previ- ously known only from the south of Central Europe, Italy, France, and from some Balkans countries. The above specimens represent the first record from Turkey. Aloconota debilicornis (Erichson, 1839) Material examined: Gümüshane: 1 ex., ca. 12km NE Gümüshane, ca. 40°31N, 39°33E, 1300 m, 12.V1.1998, leg. SOLODOVNIKOV (cAss); 3 exs., ca. 50km SW Trabzon, NE Kürtün, 40°43N, 39°13E, 1250 m, bank of stream, floated from sand and gravel, 27.VII.2006, leg. Ass- ING, SCHULKE (cAss, cSch); 1 ex., ca. 50km SW Trabzon, 9-10km S Dikkaya, ca. 40°36N, 39°29E, 2000 m, 9.V1.1998, leg. SOLODOVNIKOV (cAss). - Trabzon: 1 ex., ca. 50km S Trabzon, 20km S Macka, Altindere Milli Park, 40°40N, 39°40E, 1540 m, bank of stream, 26.VII.2006, leg. SCHULKE (cSch). — Rize: 24 exs., 30km SW Hopa, Caglayan river valley, ca. 41°15N, 41°13E, 500m, 29.V1.1998, leg. SOLODOVNIKOV (cAss, cSol); 1 ex., ca. 30km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1000 m, 26.V1.1998, leg. SOLODOVNIKOV (cAss); 4 exs., 30km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1300 m, 28.V1.1998, leg. SOLODOVNIKOV (cAss); 1 ex., ca. 30km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1800-1900 m, 26.V1.1998, leg. SOLODOVNIKOV (cAss). — Artvin: 1 ex., ca. 40km SW Artvin, Barhal river valley, ca. 40°57N, 41°29E, 1800 m, 23.V1.1998, leg. SOLODOVNIKOV (cAss). - Erzurum: 1 ex., 35km NW Tortum, Mescit Daglari, ca. 40°30N, 41°25E, 1700-2000 m, 17.V1.1998, leg. SOLODOVNIKOV (cAss). — RUSSIA: 1 ex., Krasnodar, Sochi env., Staraya Matsesta, ca. 43°34N, 39°48E, 100 m, 5.V1.1998, leg. SOLODOVNIKOV (cAss). Aloconota debilicornis was previously known neither from Turkey nor from adja- cent countries (SMETANA 2004). It is here reported from Turkey and Russia for the first time. Aloconota gregaria (Erichson, 1839) Material examined: Gümüshane: 1 ex., ca. 50km SW Trabzon, E Zigana Geg., ca. 40°37N, 39°26E, 2500 m, 11.V1.1998, leg. SOLODOVNIKOV (cAss). — Mugla: 2 exs., SE Köycegız, 36°57N, 28°44E, 10m, floodplain forest, 28.1I1.2002, leg. Asstnc (cAss). — Antalya: 3 exs., Manavgat env., 0-50 m, 2.-3.1.1991, leg. AssınG (cAss). — Mersin: 1 ex., NW Silifke, 20km W ASSING, STAPHYLINIDAE FROM TURKEY V 27: Mut, 36°36N, 33°16E, 230 m, 20.1V.2005, leg. BRACHAT & MEYBOHM (cAss). — Kahraman- maras: 1 ex, 50km W Kahramanmaras, 8km SE Andırın, 37°35N, 36°25E, 1240 m, 19.1I1.2005, leg. Assıng (cAss). — Gaziantep: 1 ex., S Birecik, 37°01N, 37°58E, 340 m, 24.1V.2004, leg. BRACHAT & MEYBOHM (cAss). The eurytopic Aloconota gregaria is one of the most widespread and common species of Aleocharinae in the Western Palaearctic region. Remarkably, it had not been reported from Turkish territory before (SMETANA 2004). Aloconota subgrandis (Brundin, 1954) Material examined: Gümüshane: 1 ex., ca. 50km SW Trabzon, 9-10km S Dikkaya, ca. 40°36N, 39°29E, 2000 m, 9.V1. 1998, leg. SOLODOVNIKOV (cAss). The species is here reported from Turkey for the first time. According to SMETANA (2004), it has not been reported from adjacent countries. Aloconota sulcifrons (Stephens, 1832) Material examined: Rize: 2 exs., 30km SW Hopa, Caglayan river valley, ca. 41°15N, 41°13E, 500 m, 29.V1.1998, leg. SOLODOVNIKOV (cAss); 2 exs., ca. 30 km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1000 m, 26.V1.1998, leg. SOLODOVNIKOV (cAss); 2 exs., ca. 30km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1800-1900 m, 26.V1.1998, leg. SOLODOVNIKOV (cAss). — Erzurum: 2 exs., 40km NW Tortum, Mescit Daglari, ca. 40°36N, 41°23E, 1800m, 21.V1.1998, leg. SOLODOVNIKOV (cAss); 1 ex., 40km NW Tortum, Mescit Daglari, ca. 40°36N, 41°23E, 2100 m, 21.V1.1998, leg. SOLODOVNIKOV (cAss); 1 ex., 30km NW Tortum, Mescit Daglari, ca. 40°40N, 41°15E, 800m, 21.V1.1998, leg. SOLODOVNIKOV (cAss). According to SMETANA (2004), this widespread and common Cosmopolitan species had not been reported from Turkey before. Amischa bifoveolata (Mannerheim, 1830) Material examined: Ordu: 4 exs., 25 km S Ordu, S Kabaduz, 40°49N, 37°54E, 990 m, mixed forest with alder, spruce, bramble, ivy, 30.VII.2006, leg. Asstnc, SCHULKE (cAss, cSch). - Artvin: 1 ex., ca. 40km SW Artvin, source of Barhal river, 41°05N, 31°30E, 2400-2800 m, 23.-24.V1.1998, leg. SOLODOVNIKOV (cAss). This widespread Palaearctic species is here recorded from Turkey for the first time. Amischa decipiens (Sharp, 1869) Material examined: Ankara: 1 ex., SE Ankara, N-Elma Dagi, 1300 m, hollow Salix trunk, 31.X.1995, leg. Vrr (cAss). According to SMETANA (2004), this species was previously unknown from Turkey. Amischa forcipata Mulsant & Rey, 1873 Material examined: Izmir: 1 ex., Bozdag, 21.V.2006, leg. Antas (cAss). The species is widespread in central and southern Europe (SMETANA 2004) and 28 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 was recently also recorded from Tunisia (AssING 2005e). This is the first record from Turkey. Ousipalia caesula (Erichson, 1839) Material examined: Mugla: 1 ex., N Fethiye, Calis, 36°41N, 29°06E, 5 m, near beach, sifted from grass roots under bushes, 1.X.2002, leg. Assıng (cAss); 1 ex., Yatagan-Bozdogan, 37°26'35N, 28°18'07E, 825m, 19.IV.2006, leg. BRACHAT & MEYBOHM (cAss). — Antalya: 2 exs., 60km SSW Antalya, Cirali, 36°25N, 30°28E, 40 m, road margin, grass and moss sifted, 23.11.2002, leg. AssınG (cAss); 1 ex., same data, but 4.IV.2002 (cAss). — Mersin: 1 ex., road from Silifke to Gülnar, ca. 15km W Silifke, 36°23N, 33°50E, 475 m, road margin, under stones, 27.X1I.2000, leg. Asstnc (cAss). - Osmaniye: 1 ex., 6km NE Osmaniye, 37°07N, 36°19E, 130 m, river bank, washed, 10.IV.2004, leg. Assıng & SCHULKE (cAss). — Kahramanmaras: 2 exs., 20km SW Hopurlu, 37°29N, 36°48E, 520 m, 27.TV.2004, leg. BRACHAT & MEYBOHM (cAss).- Antakya: 1 ex., N Belen Gecidi, 36°31N, 36°14E, 1010 m, 23.IV.2004, leg. BRACHAT & MEYBOHM (cAss). The species is widespread in the Western Palaearctic region. It is also known from Greece and Cyprus (AssING & WUNDERLE 2001, SMETANA 2004), but had not been recorded from Turkey before. Notothecta pisidica Assing, 2004 Material examined: Aydin: 1 ex., NE Aydin, Imambaba Tepesi, 37°57N, 27°53E, 1460 m, 20.1V.2006, leg. BRACHAT & MEYBOHM (cAss). — Antalya: 1 ex., Akseki, 19.111.2002, leg. Esser (cAss). Only the holotype from Konya of this recently described myrmecophile had been known (AssING 2004a). The specimen from Aydin was collected in a nest of Messor sp., suggesting that this may be the — previously unknown - host ant. Taxicera deplanata (Gravenhorst, 1802) Material examined: Trabzon: 14 exs., ca. 50 km S Trabzon, 20km S Macka, Altindere Milli Park, 40°40N, 39°40E, 1540m, bank of stream, 26.VII.2006, leg. Assıng, SCHULKE (cAss, cSch). The species is widespread in Europe, but was previously unknown from Turkey (Kapp 2005). Liogluta alpestris (Heer, 1839) Material examined: Giimiishane: 18 exs., ca. 80km SW Trabzon, Zigana Gec., 40°37N, 39°24E, 2050 m, roots of grass, moss, and shrubs sifted, 23.VII.2006, leg. AssınG (cAss). - Rize: 4 exs., 60km SSE Rize, Ovitdagi Geg., 40°38N, 40°45E, 2510m, N-slope, sifted moss and grass near rocks, 25.VII.2006, leg. AssınG (cAss). Liogluta alpestris is widespread and rather common in the Western Palaearctic re- gion and has even been recorded as far east as the Altai range, but was previously un- known from Turkey (SMETANA 2004). Liogluta microptera Thomson, 1867 Material examined: Rize: 2 exs., ca. 30km SW Hopa, Caglayan river valley, ca. 41°10N, 41°19E, 1000-1300 m, 28.V1.1998, leg. SOLODOVNIKOV (cAss). — Artvin: 1 ex., ca. 40 km SW ASSING, STAPHYLINIDAE FROM TURKEY V 29 Artvin, source of Barhal river, 41°05N, 31°30E, 2400-2800m, 23.-24.V1.1998, leg. SOLODOVNIKOV (cAss). Liogluta microptera is widespread in Europe (SMETANA 2004) and was recently even reported from China (AssınG 2005a). It is here recorded from Turkey for the first time. Atheta (Philhygra) laevigata (Hochhuth, 1849) Material examined: Mugla: 21 exs., SE Dalaman, 36°47N, 28°50E, 10 m, floodplain forest, meadows, 28.111.2002, leg. Asstnc (cAss); 1 ex., Mugla env., Bayir, 37°16N, 28°10E, 400 m, 1.V.2001, leg. MEYBOHM (cAss). — Erzurum: 1 ex., 35-40 km NW Tortum, Mescit Daglari, ca. 40°30N, 41°17E, 2600 m, 19.V1.1998, leg. SOLODOVNIKOV (cAss). — Gümüshane: 1 ex., ca. 50km SW Trabzon, 9-10km S Dikkaya, ca. 40°36N, 39°29E, 2000m, 9.V1.1998, leg. SOLODOVNIKOV (cAss). The species had become known only from Azerbaijan (SMETANA 2004). The above specimens represent the first records from Turkey. Atheta (Philhygra) lyciana Assing, 2003 Material examined: Artvin: 1 ex., ca. 40km SW Artvin, source of Barhal river, 41°05N, 31°30E, 2400-2800 m, 23.-24.V1.1998, leg. SOLODOVNIKOV (cAss). This recently described species was previously known only from the type locali- ty in Mugla province (AssING 2003a). Atheta (Philhygra) luridipennis Mannerheim, 1830 Material examined: Rize: 2 exs., 30km SW Hopa, Caglayan river valley, ca. 41°15N, 41°13E, 500 m, 29.V1.1998, leg. SOLODOVNIKOV (cAss). This widespread species was previously unknown from Turkey (SMETANA 2004). Atheta (Parameotica) epirotica Benick, 1981 (Fig. 60) Material examined: Ankara: 1 ex., SE Ankara, N-Elma Dagi, 1300 m, decaying and hollow Salix trunks, 31.X.1995, leg. Vrr (cAss). This is the first record of A. epirotica since the original description, which is based on four type specimens from Ipiros, northern Greece (BENICK 1981). The species is here reported from Turkey for the first time. For an illustration of the lateral aspect of the median lobe of the aedeagus see Fig. 60. Atheta (Microdota) speculum (Kraatz, 1856) Material examined: Izmir: 1 ex., N Izmir, Yamanlar Dagi, 38°33N, 27°09E, 940 m, grassy patch in pine forest, under stones, sifted grass roots, 28.XII.2005, leg. AssınG (cAss); 1 ex., Nif Dagı, 38°24N, 27°24E, 1010 m, 23.1V.2006, leg. BRACHAT & MEYBOHM (cAss). This apparently very rare species had become known only from Greece and Azer- baijan (SMETANA 2004). 30 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 Atheta (Microdota) subtilis (Scriba, 1866) Material examined: Trabzon: 4 exs., ca. 50km S Trabzon, 20km S Macka, Altindere Milli Park, 40°41N, 39°39E, 1650 m, spruce forest with Rhododendron, sifted, 26.VII.2006, leg. AssING, SCHULKE (cAss, cSch). — Rize: 1 ex., 50km SSE Rize, W Sivrikaya, 40°41N, 40°39E, 2050 m, natural Abies forest, litter and dead wood, 1.VIII.2006, leg. SCHULKE (cSch). Atheta subtilis is widespread in the Western Palaearctic region, but had not been reported from Turkey (SMETANA 2004). Atheta (Ceritaxa) testaceipes (Heer, 1839) Material examined: Kahramanmaras: 3 exs., 14 km SW Tirkoglu, 37°21N, 36°44E, 850 m, 19.TV.2005, leg. BRACHAT & MEYBOHM (cAss). The species is widespread in Europe and had been reported also from Cyprus (SMETANA 2004). This is the first record from Turkey. Atheta (Atheta) chefsurica (Eppelsheim, 1880) Material examined: Erzurum: 6 exs., 40km NW Tortum, Mescit Daglari, ca. 40°36N, 41°23E, 2100 m, 20.V1.1998, leg. SOLODOVNIKOV (cAss). The species was previously known only from Georgia (SMETANA 2004) and Ar- menia (AssING 2005b) and is here reported from Turkey for the first time. Atheta (Atheta) castanoptera (Mannerheim, 1840) Material examined: Ordu: 1 ex., 25 km S Ordu, S Kabaduz, 40°49N, 37°54E, 990 m, mixed forest with alder, spruce, bramble, ivy, 30.VII.2006, leg. AssınG (cAss). — Giresun: 3 exs., ca. 30km S Giresun, 40°35N, 38°27E, 1350 m, spruce forest with Rhododendron, 29.VII.2006, leg. Asstnc (cAss). — Gümüshane: 2 exs., ca. 50km SW Trabzon, NE Kürtün, 40°44N, 39°13E, 1430 m, spruce forest with Rhododendron, sifted, 27.VII.2006, leg. AssInc, SCHULKE (cAss, cSch). — Trabzon: 4 exs., ca. 50km S Trabzon, 20km S Macka, Altindere Milli Park, 40°40N, 39°40E, 1560 m, spruce forest with Rhododendron, sifted, 26. V1I1.2006, leg. AssınG (cAss); 171 exs., 20km S Macka, Altindere Milli Park, 40°41N, 39°39E, 1650 m, spruce forest with Rhododendron, leaf litter and mushrooms sifted, 26.VII.2006, leg. AssInc, SCHULKE (cAss, cSch). — Rize: 5 exs., 30km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1300 m, mushrooms, 28.V1.1998, leg. SOLODOVNIKOV (cAss, cSol); 12 exs., 50 km SSE Rize, W Sivrikaya, 40°41N, 40°39E, 2050 m, natural Abies forest, litter and dead wood, 1.VIII.2006, leg. AssING, SCHULKE (cAss, cSch); 5 exs., 30km ESE Rize, S Kaptanpasa, 40°57N, 40°47E, 690m, mixed forest with beech, alder, chestnut, spruce, Rhododendron, bramble, sifted, 5.VIII.2006, leg. AssınG (cAss). — Artvin: 1 ex., ca. 40km SW Artvin, Barhal river valley, ca. 40°57N, 41°29E, 1800 m, 23.V1.1998, leg. SOLODOVNIKOV (cAss). The species is widespread and common in the Palaearctic region, but had not been reported from Turkey before (SMETANA 2004). Atheta (Atheta) hypnorum (Kiesenwetter, 1850) Material examined: Gümüshane: 1 ex. [det. Vocer], ca. 50km SW Trabzon, NE Kürtün, 40°44N, 39°13E, 1430 m, spruce forest with Rhododendron, sifted, 27.VII.2006, leg. AssınG (cAss). The above specimen represents the first record from Turkey. ASSING, STAPHYLINIDAE FROM TURKEY V 31 Atheta (Atheta) ebenina (Mulsant & Rey, 1873) Material examined: RUSSIA: 1 ex. [det. VoGEL], Krasnodar, Sochi env., Staraya Matsesta, ca. 43°34N, 39°48E, 100 m, 5.V1.1998, leg. SOLODOVNIKOV (cAss). The species was previously unknown from the Russian South European territory (SMETANA 2004). Atheta (Tetropla) nigritula (Gravenhorst, 1802) Material examined: Gümüshane: 16 exs., ca. 50km SW Trabzon, NE Kürtün, 40°44N, 39°13E, 1430 m, spruce forest with Rhododendron, sifted, 27.VII.2006, leg. AssING, SCHULKE (cAss, cSch). — Trabzon: 2 exs., ca. 50km S Trabzon, 20km S Macka, Altindere Milli Park, 40°41N, 39°39E, 1650 m, spruce forest with Rhododendron, leaf litter and mushrooms sifted, 26.VII.2006, leg. AssınG (cAss). - Erzurum: 10 exs., 40km NW Tortum, Mescit Dagßları, ca. 40°36N, 41°23E, 2100m, 20.V1.1998, leg. SOLODOVNIKOV (cAss). — RUSSIA: 1 ex., Krasnodar, Sochi env., Staraya Matsesta, ca. 43°34N, 39°48E, 100m, 5.V1.1998, leg. SOLODOVNIKOV (cAss). These are the first records of this widespread Palaearctic species from Turkey and from the Russian South European territory (SMETANA 2004). Atheta (Datomicra) dadopora Thomson, 1867 Material examined: Giresun: 1 ex. [det. SCHULKE], ca. 30km S Giresun, 40°35N, 38°27E, 1350 m, spruce forest with Rhododendron, 29.V11.2006, leg. SCHULKE (cSch). - Giimiishane: 1 ex. [det. SCHULKE], ca. 50km SW Trabzon, NE Kürtün, 40°44N, 39°13E, 1430m, spruce forest with Rhododendron, near stream, sifted, 27.VII.2006, leg. SCHULKE (cSch). — Trabzon: 7 exs., ca. 50km S Trabzon, 20km S Macka, Altindere Milli Park, 40°41N, 39°39E, 1650 m, spruce forest with Rhododendron, leaf litter and mushrooms sifted, 26.VII.2006, leg. AssınG (cAss). — Rize: 1 ex., 30km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1300 m, mushrooms, 28.V1.1998, leg. SOLODOVNIKOV (cAss); 2 exs., 30km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1800-1900 m, 26.V1.1998, leg. SOLODOVNIKOV (cAss). — RUS- SIA: 1 ex., Krasnodar, Sochi env., Staraya Matsesta, ca. 43°34N, 39°48E, 100 m, 5.V1.1998, leg. SOLODOVNIKOV (cAss). The species is widespread in the Palaearctic region; the above specimens represent the first records from Turkey and from the Russian South European territory (SMETANA 2004). Atheta (Datomicra) dissimulans n.sp. (Figs. 50-59) Types Holotype d: TR [3] - Gümüshane, ca. 25 km SW Gümüshane, Tersundagi Geg., 2070 m, 40°17'38N, 39°18’02E, 24.VII.2006, V. Asstnc / Holotypus ¢ Atheta dissimulans sp. n. det. V. AssING 2006 (cAss). Paratypes: 74d, 22%: same data as holotype (cAss); 7 exs.: same data, but leg. M. SCHULKE (cSch). Etymology The name (Latin, adjective: deceiving, pretending) alludes to the external appearance of this species, which resembles that of a species of Ousipalia Gozis or brachypterous Microdota Mulsant & Rey rather than that of Datomicra Mulsant & Rey. 32 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 Figs. 51-60. Atheta dissimulans n.sp. (51-59) and A. epirotica (60). -— 51. Male habitus. 52. Male forebody. 53. Antenna. 54. Male sternite VIII. 55-56. Median lobe of aedeagus in lat- eral and in ventral view. 57. Apical lobe of paramere. 58. Female sternite VIII. 59. Spermathe- ca. 60. Median lobe of aedeagus in lateral view. — Scale bars: 1 mm (51), 0.5 mm (52-53), 0.2 mm (54, 58), 0.1 mm (55-57, 59-60). Description TL: 2.3-2.9mm. Habitus as in Fig.51. Coloration: body blackish, with the ab- dominal apex (segments VIII and following) and sometimes the pronotum and the elytra indistinctly paler blackish brown; legs yellowish; antennae blackish, with the basal 2-3 antennomeres slightly paler, dark brown. Head 1.05-1.10 times as wide as long, widest behind eyes, and with sexual dimor- phism; surface with distinct microreticulation and only with weak shine; punctura- tion sparse and extremely fine, barely noticeable; eyes rather small and weakly prominent, postocular region approximately 1.5 times the length of eyes in dorsal view (Fig.52). Antennae gradually incrassate apically; antennomere III short, less than 1.5 times as long as wide; IV transverse; V-X of increasing width and increas- ingly transverse; X approximately 1.5 times as wide as long; XI approximately as long as the combined length of IX-X (Fig. 53). Pronotum relatively large, approximately 1.2 times as wide as long and 1.25-1.30 times as wide as head, maximal width in anterior half; posterior angles rounded, weakly marked, almost obsolete; with sexual dimorphism (Fig. 52); surface with dis- ASSING, STAPHYLINIDAE FROM TURKEY V 33 tinct microreticulation and only subdued shine; puncturation very fine and moder- ately dense; pubescence fine, directed cephalad along midline and almost transverse- ly laterad in lateral areas. Elytra about 1.1 times as wide and at suture approximately 0.75 times as long as pronotum (Fig.52); puncturation very fine, slightly denser than that of pronotum; microreticulation distinct. Hind wings reduced. Abdomen approximately as wide as elytra; puncturation moderately dense and more distinct than that of forebody, distinctly sparser on tergite VII than on anteri- or tergites; posterior margin of tergite VII with palisade fringe; tergite VIII without distinct sexual dimorphism, posterior margin in both sexes weakly convex. 3: head extensively flattened or shallowly impressed; pronotum along middle with broad depression or shallow impression (Fig. 52); sternite VIII distinctly longer and less transverse than tergite VIII, its posterior margin strongly convex (Fig. 54); median lobe of aedeagus of distinctive shape (Figs. 55-56); apical lobe of paramere of similar morphology and chaetotaxy as in other species of the subgenus (Fig. 57). ?: head weakly convex in cross-section; median impression of pronotum con- fined to posterior half and much shallower and narrower; sternite VIII distinctly transverse, its posterior margin broadly convex and in the middle indistinctly con- cave (Fig. 58); spermatheca as in Fig. 59. Comparative notes From other Western Palaearctic Datomicra species, A. dissimulans is easily sepa- rated by the distinctive genitalia, as well as by conspicuous external characters, espe- cially the large pronotum, the sexual dimorphism of head and pronotum, the short elytra, the reduced hind wings, and the absence of a dimorphism of the abdominal tergite VIII. The new species in fact much more resembles species of the subgenus Microdota or of the genus Ousipalia than its consubgeners, which are generally winged, often distinctly smaller, and inhabitants of short-lived habitats (rotting or- ganic matter like mushrooms, excrements, compost, etc.). Distribution and bionomics The type locality is situated about 25km to the southwest of Gümüshane, Gümüshane province (Fig.50), at an altitude of 2070 m. It is illustrated in AssınG (2006i: fig. 85). The specimens were collected by sifting the litter of a montane spruce forest, suggesting that this species is an inhabitant of the forest floor. Atheta (Dimetrota) aeneipennis (Thomson, 1856) Material examined: Trabzon: 1 ex., ca. 50km S Of, S Uzungöl, 40°36N, 40°17E, 2050 m, moss and roots of grass and herbs near rocks, sifted, 4.VIII.2006, leg. AssınG (cAss); 1 ex., ca. 50km S Of, S Uzungöl, 40°36N, 40°18E, 1870m, spruce forest, 4.VIII.2006, leg. Asstnc (cAss). - Rize: 8 exs., 30km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1300 m, mushrooms, 28.V1.1998, leg. SOLODOVNIKOV (cAss, cSol); 17 exs., 30 km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1800-1900 m, 26.V1.1998, leg. SOLODOVNIKOV (cAss, cSol). This species has become known from most of the Palaearctic region and also from the Oriental region (SMETANA 2004). It is here reported from Turkey for the first time. 34 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 Atheta (Dimetrota) knabli Benick, 1938 Material examined: Giimiishane: 18 exs., ca. 80km SW Trabzon, Zigana pass, 40°37N, 39°24E, 2050 m, roots of grass, moss, and shrubs sifted, 23.VII.2006, leg. AssınG (cAss). - Trabzon: 4 exs., ca. 50 km S Of, S Uzungöl, 40°36N, 40°17E, 2050 m, moss and roots of grass and herbs near rocks, sifted, 4.VIII.2006, leg. AssınG (cAss); 1 ex., ca. 50 km S Of, S Uzungöl, 40°36N, 40°18E, 1870 m, spruce forest, 4.VIII.2006, leg. AssınG (cAss). Previously published records are confined to the Alps (SMETANA 2004), but ac- cording to J. VOGEL (pers. comm.) the species was also collected in Bulgaria. It is here reported from Turkey for the first time. Atheta (Dimetrota) cinnamoptera (Thomson, 1856) Material examined: Rize: 3 exs., 30km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1300 m, mushrooms, 28.V1.1998, leg. SOLODOVNIKOV (cAss, cSol); 7 exs., 30 km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1800-1900m, 26.V1.1998, leg. SOLODOVNIKOV (cAss, cSol). — Artvin: 1 ex., ca. 40km SW Artvin, Barhal river valley, ca. 40°57N, 41°29E, 1800 m, 23.V1.1998, leg. SOLODOVNIKOV (cAss). These are the first records of this widespread Palaearctic species from Turkey. Atheta (Dimetrota) episcopalis Bernhauer, 1910 Material examined: Erzurum: 1 ex. [det. VOGEL], 30km NW Tortum, Mescit Daglani, ca. 40°40N, 41°15E, 800m, 21.V1.1998, leg. SOLODOVNIKOV (cAss); 1 ex. [det. Vocer], 40km NW Tortum, Mescit Daglari, ca. 40°36N, 41°23E, 2100 m, 20.V1.1998, leg. SOLODOVNIKOV (cAss). The species is here recorded from Turkey for the first time. Atheta (Dimetrota) laevana (Mulsant & Rey, 1852) Material examined: Trabzon: 1 ex., 20km S Macka, Altindere Milli Park, 40°41N, 39°39E, 1650 m, spruce forest with Rhododendron, leaf litter and mushrooms sifted, 26.VII.2006, leg. AssınG (cAss). The above specimen represents the first record of the species from Turkey. Atheta mucronata (Kraatz, 1859) Material examined: Antalya: 7 exs., Manavgat env., Kizilot, 0-50 m, 2.1.1991, leg. AssInG (cAss). In the Palaearctic catalogue (SMETANA 2004), this name is erroneously listed as a junior synonym of Pelioptera opaca (Kraatz, 1857). The species is widespread in the Mediterranean region (FELDMANN in prep.). It is here reported from Turkey for the first time. Atheta sodalis (Erichson, 1837) Material examined: Giresun: 1 ex., Giresun, ca. 30 km S Giresun, 40°36N, 38°27E, 1250 m, spruce forest with Rhododendron, sifted, 29.VII.2006, leg. SCHULKE (cSch). - Gümüshane: 6 exs., ca. 25km SW Gümüshane, Tersundagi Gec., 40°18N, 39°18E, 2070 m, N-slope, spruce forest, litter and dead wood sifted, 24.VII.2006, leg. AssING, SCHULKE (cAss, cSch). — Trab- zon: 4 exs., ca. 50km S Trabzon, 20km S Macka, Altindere Milli Park, 40°40N, 39°40E, ASSING, STAPHYLINIDAE FROM TURKEY V 39 1560 m, spruce forest with Rhododendron, sifted, 26.VII.2006, leg. Assıng, SCHULKE (cAss, cSch); 2 exs., 20km S Macka, Altindere Milli Park, 40°41N, 39°39E, 1650 m, spruce forest with Rhododendron, leaf litter and mushrooms sifted, 26.VII.2006, leg. Assinc, SCHULKE (cAss, cSch). The species was previously unknown from Turkey (SMETANA 2004). Atheta britanniae Bernhauer & Scheerpeltz, 1926 Material examined: Giimiishane: 2 exs. [det. SCHULKE], ca. 25 km SW Gümüshane, Ter- sundagi Gec., 40°18N, 39°18E, 2070m, N-slope, spruce forest, litter and dead wood sifted, 24.V11.2006, leg. SCHULKE (cSch). - Trabzon: 26 exs. [det. VOGEL, SCHULKE], 20 km S Macka, Altindere Milli Park, 40°41N, 39°39E, 1650 m, spruce forest with Rhododendron, leaf litter and mushrooms sifted, 26.VII.2006, leg. Assıng, SCHULKE (cAss, cSch). — RUSSIA: 1 ex. [det. VOGEL], Krasnodar, Sochi env., Staraya Matsesta, ca. 43°34N, 39°48E, 100 m, 5.V1.1998, leg. SOLODOVNIKOV (cAss). The species was previously unknown from Turkey and from the Russian South European territory (SMETANA 2004). Atheta circassica Bernhauer, 1900 Material examined: Rize: 3 exs., 30km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1300m, mushrooms, 28.V1.1998, leg. SOLODOVNIKOV (cAss). — RUSSIA: 1 ex. [det. VOGEL], Krasnodar, Sochi env., Staraya Matsesta, ca. 43°34N, 39°48E, 100 m, 5.V1.1998, leg. SOLODOVNIKOV (cAss). Atheta circassica had become known only from the West Caucasus region and is here reported from Turkey for the first time. Atheta fungicola (Thomson, 1852) Material examined: Rize: 4 exs., 30km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1300 m, mushrooms, 28.V1.1998, leg. SOLODOVNIKOV (cAss, cSol). The known distribution of this species includes most of Europe. It is here report- ed from Turkey for the first time. Atheta gagatina (Baudi di Selve, 1848) Material examined: Erzurum: 4 exs., 40km NW Tortum, Mescit Daglari, ca. 40°36N, 41°23E, 2100 m, 20.V1.1998, leg. SOLODOVNIKOV (cAss, cSol). The above specimens represent the first record of this common Palaearctic species from Turkey. Acrotona muscorum (Brisout, 1860) Material examined: Izmir: 3 exs., Boz Daglar, above Bozdag, road to ski resort, 38°21N, 28°06E, 1480m, N-slope with grass and stones, sifted, 3.1V.2006, leg. Assinc, WUNDERLE (cAss, cWun); 22 exs., Boz Daglar, above Bozdag, road to ski resort, 38°21N, 28°07E, 1500 m, N-slope, Alnus and Salix litter sifted, 3.TV.2006, leg. Assıng, WUNDERLE (c Wun). — Erzurum: 3 exs., 40km NW Tortum, Mescit Daglari, ca. 40°36N, 41°23E, 2100m, 20.V1.1998, leg. SOLODOVNIKOV (cAss). 36 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 This widespread species was recently recorded from Turkey for the first time (AssING 2006f). Acrotona nigerrima (Aubé, 1850) Material examined: Izmir: 1 ex., Nif Dagi, 38°24N, 27°24E, 970m, 24.IV.2006, leg. BRACHAT & MEYBOHM (cAss). According to SMETANA (2004), this widespread Western Palaearctic species was unknown from Turkey. Acrotona piceorufa (Mulsant & Rey, 1873) Material examined: Isparta: 3 exs., Egridir env., Kovada Gölü, 37°37N, 30°51E, 1000 m, 13.V.2000, leg. MEYBOHM (cAss). — Antalya: 1 ex., W Antalya, Saklikent, 1000 m, 11.V.2000, leg. MEYBOHM & Bracuat (cAss). — Mersin: 1 ex., Kirobası-Güzeloluk, 36°45N, 33°58E, 1430 m, 7.-8.V.2004, leg. BESUCHET (cAss). In Turkey, A. piceorufa was previously known only from Osmaniye (AssınG 2006). Drusilla limata Assing, 2005 Material examined: Antalya: 1 ex., Korkuteli, Yazır, 1.V.1990, leg. GILLERFORS (cWun). This recently described species was previously known only from two localities in Antalya (AssING 2005c). Drusilla lydica n.sp. (Figs. 61-71) Types Holotype 2: TR [6] - Aydin, 15km NNE Aydin, Imambaba T., 1420m, u. stones, 37°56'53N, 27°53'45E, 5.1V.2006, V. Asstnc / Holotypus 2 Drusilla lydica sp. n. det. V. Ass- ING 2006 (cAss). Paratypes: 12 dd, 2 22: same data as holotype (cAss); 20 exs.: same data, but leg. Wun- DERLE (cWun); 2 22: TR [5] — Aydin, 15km NNE Aydin, Imambaba T., 1410 m, 37°56'38N, 27°53’40E, 5.1V.2006, V. AssınG (cAss); 5 exs.: same data, but leg. WUNDERLE (cWun); 9 dd, 2 2¢: TR [8] - Aydin, 15km NNE Aydin, Imambaba T., below peak, 1600 m, 37°57'16N, 27°53'55E, 5.1V.2006, V. Asstnc (cAss); 18,4 22: N37°56'47 E027°53'53 (8), Türkei, Aydin, Pasayaylasi, 1460 m, 20.TV.2006, |. BRACHAT & MEYBOHM (cAss); 18,3 22: TR Prov.: Aydin (8), N Aydin, Pasayaylasi, 1460 m, 20.IV.2006, N37°56'47", E27°53'53”, leg. MEYBOHM & BRACHAT (cAss); 2 22: TR Prov.: Aydin (9), N Aydin, Str. -> Pasayaylasi, 1115 m, 20.1V.2006, N37°55'46", E27°53'46", leg. MEYBOHM & BRACHAT (cAss); 18,2 22: TR [17] - Izmir, 5 km S Tire, N-slope with stones and shrubs, 990 m, 38°03’04N, 27°45’07E, 9.IV.2006, V. AssınG (cAss); 3 exs.: same data, but leg. WUNDERLE (cWun). Etymology The name (adjective) is derived from Lydia, the ancient name of the region where the species was found. Description TL: 4.2-5.1 mm. Habitus as in Fig. 61. Coloration: body blackish, with the elytra often slightly to distinctly paler, brown to dark brown; legs pale yellowish brown; antennae dark brown with the basal 2-3 antennomeres yellowish to reddish. ASSING, STAPHYLINIDAE FROM TURKEY V 37 Figs. 61-70. Drusilla lydica n. sp. - 61. Male habitus. 62. Male forebody. 63. Female forebody. 64. Antenna. 65. Male tergite VIII. 66. Median lobe of aedeagus in lateral view. 67. Female ter- gite VIII. 68-70. Spermathecae of three females in different aspects. — Scale bars: 1mm (61-63), 0.5 mm (64), 0.2 mm (65-67), 0.1 mm (68-70). Head approximately as wide as long or weakly oblong; eyes relatively small, but projecting from lateral outline of head, postocular region nearly twice as long as eyes in dorsal view; dorsal surface without distinct microsculpture; puncturation fine and moderately sparse, but distinct (Figs. 62-63). Antenna as in Fig. 64. Pronotum with weakly pronounced sexual dimorphism (Figs. 62-63); slender, 1.10-1.15 times as long as wide and approximately 1.1 times as wide as head; punc- turation very distinct, somewhat granulose, and dense to very dense; interstices without microsculpture. Elytra approximately 1.25 times as wide and at suture 0.60-0.65 times as long as pronotum; laterally with shallow longitudinal impressions (Figs. 62-63); punctura- 38 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 tion very coarse, coarser than that of pronotum, dense, and granulose; interstices without microsculpture. Hind wings reduced. Metatarsomere I approximately as long as the combined length of II-III. Abdomen about 1.25 times as wide as elytra, widest at segment V; puncturation fine and moderately sparse, distinctly denser on anterior than on posterior tergites; anterior impressions of tergites III-V deep; tergite VII with very narrow rudiment of a palisade fringe; microsculpture absent. 3: pronotum with distinct impression along median line (Fig. 62); posterior mar- gin of tergite VIII broadly concave and serrate (Fig. 65); median lobe of aedeagus as in Fig. 66. ?: pronotum with shallower and narrower impression; posterior margin of ter- gite VIII serrate, less broadly concave than in d (Fig. 67); posterior margin of stern- ite VIII broadly convex, not concave in the middle; spermatheca of distinctive shape (Figs. 68-70). Comparative notes In a recent revision eight species were recorded from Turkey, the widespread D. canaliculata (Fabricius) and seven species with restricted distributions, the latter oc- curring in southern Anatolia from western Antalya to Gaziantep and Mardin (Ass- ING 2005c). In order to accommodate the new species, the key provided in this revi- sion is modified as follows: 18 Pronotum densely and granulosely punctured (Figs. 62-63). Spermatheca with trans- versely dilated capsule (Figs. 68-70). — Western Aydin Dagları (Aydin, Izmir) (Fig. 71). . Era oe or ee ee EN aan. AR ee ee ree eee Pe D. lydica n. sp. — Pronotum less densely and more finely punctured. Spermatheca with capsule of com- pletely different shape. — Species from southern Anatolia. ............ 0.00000 eee 18a 18a Elytra relatively long, at suture approximately 0.65 times as long as pronotum (AssING 2005c: fig. 82). Spermatheca as in AssiNG (2005c: fig. 87). ........... D. cernens Assing — Elytra at suture at most about 0.60 times as long as pronotum. Spermatheca of different Shape man kon Wa oar, MLE, vr EL comets Sige ER he RE an hme > I Baer a 19 Distribution and bionomics All the type specimens were found in several localities in the western Aydin Dagları (Fig. 71). It seems surprising that the species had not been discovered before, since it is apparently not uncommon in this mountain range. The types were collect- ed under stones in grassy vegetation close to — and in the periphery of - pine forests and in a pasture at altitudes of 990-1600 m. An association with particular ant species was not observed. Myrmoecia plicata (Erichson, 1837) Material examined: Manisa: 1 ex., ca. 10km S Manisa, Karadag, 38°33N, 27°25E, 1270 m, N-slope, pasture with stones, under stone, 15.1V.2006, leg. AsstnG (cAss). — Kastamonu: 1 ex., Ilgazdagi Geg., 1700 m, 28.VIII.1988, leg. RreEDEL (cWun). The species was only recently reported from Turkey for the first time (AssING 2006). ASSING, STAPHYLINIDAE FROM TURKEY V 32 Fig. 71. Distributions of Dinusa smyrnensis n. sp. (@), Drusilla lydica n. sp. (©), Megalogastria cingulata (Eppelsheim) (0), Medon reliquus n.sp. (@), Pyroglossa pontica n.sp. (MM), and Cousya schuelkei n. sp. (MM). Myrmoecia rigida (Erichson, 1839) Material examined: Mardin: 1 ex., Midyat, 30.V.1987, leg. SCHONMANN & SCHILLHAMMER (NHMW). This is the first record of M. rigida from Turkey. Pella hampei (Kraatz, 1862) Material examined: Izmir: 1 ex., ca. 25km NE Aydin, WSW Hamamköy, Murtat Dagi, 1230 m, 38°01N, 27°57E, 9.TV.2006, leg. AsstNc (cAss). This rare species had been known only from the southeast of Central Europe and from southeastern Europe (MARUYAMA 2006); it is here reported from Turkey for the first time. Pella laeviceps (Eppelsheim, 1880) Material examined: Gümüshane: 1 ex., ca. 12km NE Gümüshane, ca. 40°31N, 39°33E, 1300 m, 12.V1.1998, leg. SOLODOVNIKOV (cAss). — Trabzon: 3 exs., ca. 50km S Ardesen, Cat, 40°52N, 40°56E, 1240 m, alder forest, sifted, 3. VIII.2006, leg. Assıng, SCHULKE (cAss, cSch). Previous records of this species are confined to the Caucasus region (Russia, Georgia) (MARUYAMA 2006); it is here reported from Turkey for the first time. Pella laticollis (Markel, 1845) Material examined: Sinop: 2 exs., Cangal Dagi, 8.-16.VII.1961, leg. SCHUBERT (NHMW, cAss). According to Maruyama (2006), this species was previously unknown from Turkey. 40 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 Pella lugens (Gravenhorst, 1802) Material examined: Izmir: 2 exs., ca. 25km NE Aydin, WSW Hamamköy, Murtat Dagi, 1230 m, 38°01N, 27°57E, 9.1V.2006, leg. Assıng, WUNDERLE (cAss, cWun). The known distribution of P lugens ranges from western Europe to Middle Asia (MARUYAMA 2006, SMETANA 2004), but there had been no previous records from Turkey. Pella ruficollis (Grimm, 1845) Material examined: Izmir: 69 exs., ca. 25 km NE Aydin, WSW Hamamköy, Murtat Dagi, 1230 m, 38°01N, 27°57E, 9.TV.2006, leg. Assıng, WUNDERLE (cAss, cFel, cWun). MARUYAMA (2006) reports this species for Turkey, but does not specify any local- ities. Pella gibbera n.sp. (Figs. 72-83) Types Holotype d: TR [18] - Izmir, WSW Hamamköy, Murtat Dagi, 1230m, 38°00'52N, 27°56'56E, 9.1V.2006, V. Assinc / Holotypus 6 Pella gibbera sp. n. det. V. Asstnc 2006 (cAss). Paratypes:2 4d, 3 29: same data as holotype (cAss); 1 2: same data, but leg. WUNDERLE (cWun). Etymology The name (Latin, adjective: humpy) refers to the distinctive protuberance of the sperma- thecal duct. Description Measurements (inmm) and ratios (range; n = 6): AL: 1.86-1.92; HL: 0.72-0.77; HW: 0.86-0.91; PW: 1.10-1.19; PL: 0.79-0.86; EL: 0.72-0.79; EW: 1.40-1.51; AW: 1.30-1.46; TiL: 0.92-0.98; TaL: 0.72-0.77; ML: 1.15-1.22; TL: 5.6-6.5; HL/HW: 0.83-0.86; PW/HW: 1.28-1.32; PW/PL: 1.38-1.42; EL/PL: 0.84-1.00; EW/PW: 1.23-1.30; AW/EW: 0.92-0.98; TiL/TaL: 1.22-1.29. Habitus as in Fig. 72. Coloration: head castaneous brown, in most specimens dif- fusely darkened in median dorsal area; pronotum dark brown with the lateral mar- gins broadly and the anterior and posterior margin narrowly paler, castaneous brown; elytra yellowish to yellowish brown, with the scutellar area and the posteri- or 2/3-3/4 of lateral margins infuscate; abdomen blackish, with the posterior margins of the segments yellowish to yellowish brown; legs reddish brown to castaneous; an- tennae brown, with the basal 3 antennomeres reddish to reddish brown. Head moderately transverse; puncturation very fine and moderately dense; mi- crosculpture present, but shallow; eyes prominent, approximately as long as postoc- ular region in dorsal view (Fig. 73). Antenna as in Fig. 74; antennomeres III-X1 flat- tened, flatly oval in cross-section; antennomere XI approximately as long as the combined length of VIII-X. Pronotum distinctly transverse and somewhat wider than head (see measurements and ratios PW/PL and PW/HW); maximal width in anterior half, more strongly ta- pering caudad than cephalad; posterior angles weakly marked or rounded; lateral ASSING, STAPHYLINIDAE FROM TURKEY V 41 Figs. 72-82. Pella gibbera n.sp. — 72. Habitus. 73. Forebody. 74. Antenna. 75. Male tergite VII. 76. Male sternite VIII. 77-78. Median lobe of aedeagus in lateral and in ventral view. 79. Female tergite VIII. 80. Female sternite VIII. 81-82. Spermatheca. — Scale bars: 1mm (72-73), 0.5 mm (74-80), 0.2 mm (81-82). margins each with several longer setae; pronotal hypomera distinctly visible in later- al view; puncturation as fine as that of head, but somewhat denser; microsculpture absent or extremely shallow (Fig. 73). Elytra wider and at suture as long as or slightly shorter than pronotum (see mea- surements and ratios); posterior margin near posterior angles weakly sinuate; punc- turation and pubescence very fine and dense; microsculpture shallow. Hind wings fully developed. Metatarsomere I longer than the combined length of II-III, but shorter than the combined length of I-IV. Abdomen almost as wide as elytra, widest at segments V-V]; fine puncturation on tergites IV-VII present only in posterior half, pubescence on tergites VI-VII present only at posterior margin, on tergites IV-V more extensively extending cephalad in 42 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 Fig. 83. Distributions of Pella gibbera n. sp. (@), Astenus occiduus n. sp. (©), Pseudocalea mes- sorphila n. sp. (M), and Oxypoda subspectabilis n. sp. (0). lateral areas; all tergites with fine transverse microsculpture; posterior margin of ter- gite VII with palisade fringe. 3: tergite VIII transverse, its posterior margin weakly convex, in the middle trun- cate to indistinctly concave (Fig. 75); sternite VIII distinctly longer than tergite VIII and less transverse, its posterior margin strongly convex (Fig.76); median lobe of aedeagus with slender and apically acute ventral process (Figs. 77-78). 2: posterior margin of tergite VIII in the middle distinctly concave (Fig. 79); ster- nite VIII strongly transverse, shorter than that of d, posterior margin broadly con- vex (Fig. 80); spermatheca of distinctive morphology, duct with pronounced protu- berance (Figs. 81-82). Comparative notes The Palaearctic representatives of the genus were recently revised by MaRUYAMA (2006). Attributing P gibbera to any of the species groups defined in this revision proves difficult. The new species is highly distinctive and readily separated from all its Palaearctic congeners not only based on the characteristic morphology of the pri- mary sexual characters, but also by its conspicuous coloration, the shape of the an- tennae, as well as by the shapes of the male and female tergites VIII. Distribution and bionomics The type locality is situated in the western Aydin Dagları (Fig. 83). The specimens were collected by sifting oak leaf litter and grass roots on a grassy north slope with numerous rocks and scattered old oak trees at an altitude of 1230m. They were found together with P hampei, P. lugens, and numerous specimens of P. ruficollis. Liometopum microcephalum (Panzer), according to the literature the host ant of P hampei and P. ruficollis, was not seen in the field and absent from the sifted soil sam- ples. ASSING, STAPHYLINIDAE FROM TURKEY V 43 Pyroglossa pontica n.sp. (Figs. 71, 84-97) Type Holotype d [most of right antenna missing]: TR [5] — Rize, ca. 50 km SSE Rize, Ovitdagi Geg., 2510m, under stones, 40°37'31N, 40°45’27E, 25.VII.2006, V. Asstnc / Holotypus 3 Pyroglossa pontica sp. n. det. V. AssınG 2006 (cAss). Etymology The name (adjective) is derived from Pontus, the ancient name of the region where the type locality is situated. Description Measurements (inmm) and ratios: AL: 1.49; HL: 0.56; HW: 0.56; PW: 0.63; PL: 0.60; EL: 0.60; EW: 0.95; AW: 0.80; TiL: 0.76; TaL: 0.60; ML: 0.62; TL: 4.5; HL/HW: 1.00; PW/HW: 1.14; PW/PL: 1.05; EL/PL: 1.00; EW/PW: 1.50; AW/EW: 0.84; lik flab: 1,25. Habitus as in Fig. 84. Coloration: head and abdomen black; pronotum and elytra blackish brown; legs pale brown; antennae brown, gradually becoming darker to- wards apex. Head as wide as long (Fig. 85); puncturation very shallow, ill-defined, moderately dense, interstices on average approximately as wide as diameter of punctures; in- tegument with shallow fine microreticulation; pubescence relatively short, fine, and suberect; eyes moderately large (Fig. 87) and weakly projecting from lateral outline of head, somewhat shorter than postocular region in dorsal view. Maxillary palpus with third joint slender, almost 3 times as long as wide, almost twice as long as fourth joint (Fig. 88). Ligula long and slender, deeply bifid (Fig. 89). Labrum as in Fig. 90. Antenna slender; antennomere III approximately as long as II; IV oblong, distinctly shorter than III; V-X gradually increasing in width, oblong; X slightly longer than wide; XI of ovoid shape, shorter than the combined length of IX-X (Fig. 86). Pronotum very weakly transverse (see measurements and ratios PW/HW and PW/PL), maximal width in anterior half; posterior angles rounded, ill-defined; puncturation coarser and denser than that of head, interstices on average narrower than diameter of punctures; microsculpture slightly more distinct than that of head; pubescence similar to that of head, depressed, directed cephalad along midline, and predominantly laterad in lateral areas (Fig. 85). Elytra distinctly wider than and at suture approximately as long as pronotum (see ratio EL/PL and Fig. 85); posterior margin sinuate near posterior angles; punctura- tion more distinct than that of pronotum; microsculpture similar to that of prono- tum; pubescence fine, predominantly directed diagonally latero-caudad. Hind wings present and fully developed. Metatarsus somewhat shorter than metatibia (see ratio TiL/TaL); metatarsomere I as long as the combined length of I-IV. Abdomen subparallel (Fig. 91); anterior impressions of tergites III-V distinct, that of tergite VI shallow; puncturation fine, distinct, and rather sparse, denser on ante- rior than on posterior tergites; tergites with very shallow microsculpture; posterior margin of tergite VII with palisade fringe; posterior margin of tergite VIII convex, in the middle shortly truncate (Fig. 92). 3: posterior margin of sternite VIII pointed (Fig. 93); median lobe of aedeagus of distinctive shape, rather massive, and with distinctly carinate ventral process (Figs. 94-96); paramere as in Fig. 97. 2: unknown. 44 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 Figs. 84-97. Pyroglossa pontica n. sp. — 84. Habitus. 85. Forebody. 86. Antenna. 87. Head in lateral view. 88. Maxillary palpus. 89. Labium. 90. Labrum. 91. Abdomen. 92. Male tergite VIII. 93. Male sternite VIII. 94-96. Median lobe of aedeagus in lateral and in ventral view. 97. Paramere. — Scale bars: 1mm (84), 0.5mm (85-87, 91-93), 0.2mm (88, 94-97), 0.1 mm (89-90). ASSING, STAPHYLINIDAE FROM TURKEY V 45 Comparative notes The genus has a Holarctic distribution with two species known from the Nearc- tic (AssING 1999) and four species from the Palaearctic (AssING & WUNDERLE 1997, SMETANA 2004). Only one species has become known from the Western Palaearctic region, Pyroglossa pulcherrima (Bernhauer), which is distributed from eastern Siberia and Mongolia to Scandinavia. In northern Europe only females of P pulcher- rima have been recorded, suggesting that it is parthenogenetic at least in the north- west of its range (AssING & WUNDERLE 1997). Pyroglossa pontica is readily distin- guished from P pulcherrima by much smaller size, distinctly more slender antennae, much finer and sparser puncturation of the whole body, more shiny appearance, less slender head, ill-defined posterior angles of the pronotum, shorter elytra, a posteri- orly more distinctly tapering abdomen, the absence of a pronounced anterior im- pression on tergite VI, the almost impunctate anterior impressions on the anterior abdominal tergites, and the much longer metatarsomere I (in P pulcherrima as long as the combined length of II-III at most). Distribution and bionomics The type locality is situated at the Ovitdagı Gegidi (Rize), one of the major pass- es in the western parts of the Dogu Karadeniz Dagları (Fig. 71). The holotype was discovered under a stone in regularly grazed grassland at an altitude of little more than 2500 m. Cousya nigrata (Fairmaire & Laboulbene, 1856) Material examined: Aydin: 1 ex., ca. 22km NE Kuyucak, Bayrak Tepe, 37°58N, 28°34E, 900 m, N-slope, oak litter and grass sifted, 7.[V.2006, leg. AssınG (cAss). — Mugla: 2 exs., ca. 20km SW Mugla, N Meke, 37°13N, 28°12E, 590 m, pasture with stones at roadside, sifted and under stones, 12.1V.2006, leg. Assıng (cAss). — Mersin: 1 ex., 33km N Silifke, 36°36N, 33°54E, 1270 m, 18.1V.2005, leg. BRACHAT & MEYBOHM (cAss). - Antakya: 1 ex., 10km S Is- kenderun, 36°29N, 36°09E, 760 m, 4.1V.2004, leg. AssınG (cAss); 2 exs., 9km SE Iskenderun, 6km NE Belen, 36°32N, 36°15E, 1480 m, snowfield, 4.1V.2004, leg. AssınG (cAss). - Kahra- manmaras: 1 ex., 50km W Kahramanmaras, 8km SE Andırın, 37°33N, 36°26E, 1110m, 19.111.2005, leg. AssınG (cAss). - Gaziantep: 2 exs., 25km WNW Gaziantep, Kartal Dagı, 37°11N, 37°08E, 1070 m, 9.1V.2004, leg. AssınG (cAss); 3 exs., 33km E Osmaniye, NE Nurdagi Gec., 37°08N, 36°37E, 1520 m, 2.IV.2004, leg. AssınG (cAss). - Adıyaman: 1 ex., 50km NE Adıyaman, 8km NE Narince, 37°55N, 38°49E, 870 m, 24.111.2005, leg. Asstnc (cAss). According to SMETANA (2004), this species was previously unknown from Turkey. Cousya schuelkei n.sp. (Figs. 71, 98-109) Types Holotype d: TR [5a] - Rize, ca. 50km SSE Rize, Ovitdagi Geg., 2510m, sifted, 40°37'31N, 40°45'27E, 25.VII.2006, M. ScHULKE / Holotypus d Cousya schuelkei sp. n. det. V. AssING 2006 (cAss). Paratypes:2 6,1 [14 teneral]: same data as holotype (cSch, cAss). Etymology The species is dedicated to my colleague and friend MICHAEL SCHULKE, who collected the type series during a joint field trip to northeastern Anatolia. 46 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 H 105 Figs. 98-109. Cousya schuelkei n. sp. — 98. Habitus. 99. Forebody. 100. Antenna. 101. Head in lateral view. 102. Abdomen. 103. Male sternite VIII. 104. Median lobe of aedeagus in lateral view. 105. Apex of median lobe in ventral view. 106. Apical lobe of paramere. 107. Female ter- gite VIII. 108. Female sternite VIII. 109. Spermatheca. — Scale bars: 1mm (98), 0.5 mm (99, 101-102), 0.2 mm (100, 103, 107-108), 0.1 mm (104-106, 109). Description Measurements (inmm) and ratios (range; n = 4): AL: 0.82-0.88; HL: 0.41-0.42; HW: 0.39-0.41; PW: 0.47-0.48; PL: 0.39-0.42; EL: 0.38-0.41; EW: 0.60-0.62; AW: 0.53-0.54; TiL: 0.39-0.40; TaL: 0.32-0.33; ML: 0.42-0.44; TL: 2.9-3.2; HL/HW: 1.00-1.08; PW/HW: 1.19; PW/PL: 1.15-1.19; EL/PL: 0.96-0.98; EW/PW: 1.28-1.29; AW/EW: 0.88-0.90; TiL/TaL: 1.20-1.24. Habitus as in Fig. 98. Coloration: body blackish with the abdominal apex (seg- ments VIII and following) slightly paler; legs dark brown; antennae blackish. Head approximately as wide as long or weakly oblong; puncturation moderately dense and extremely fine, barely noticeable; integument with distinct fine mi- ASSING, STAPHYLINIDAE FROM TURKEY V 47 croreticulation and only with subdued shine (Fig.99); eyes moderately large (Fig. 101) and weakly projecting from lateral outline of head, slightly shorter than postocular region in dorsal view. Antenna with antennomeres I and II of subequal length; III slightly shorter than II; IV weakly transverse; V-X of gradually increas- ing width and increasingly transverse; X approximately twice as wide as long; XI about as long as the combined length of IX-X (Fig. 100). Third joint of maxillary palpus approximately 3 times as long as wide. Ligula distinctly bifid. Pronotum moderately transverse and wider than head (see measurements and ra- tios PW/PL and PW/HW); maximal width approximately in the middle, more strongly tapering cephalad than caudad; posterior angles weakly marked, almost rounded; pronotal hypomera visible in lateral view; puncturation similar to that of head; microreticulation more pronounced than that of head; pubescence directed caudad along midline and almost transversely laterad in lateral areas (Fig. 99). Elytra distinctly wider than and at suture almost as long as pronotum (see mea- surements and ratios); posterior margin near posterior angles sinuate; puncturation much more distinct than that of head and pronotum and dense, interstices narrower than diameter of punctures (Fig. 99). Hind wings fully developed. Metatarsomere I approximately as long as the combined length of H-IV. Abdomen subparallel, narrower than elytra (see ratio AW/EW); tergites III-V with anterior impressions; puncturation very fine and moderately dense; all tergites with pronounced microreticulation and only with subdued shine (Fig. 102); posteri- or margin of tergite VII with narrow rudiment of a palisade fringe; tergite VIII with- out distinct sexual dimorphism, posterior margin convex, in the middle sometimes indistinctly concave (Fig. 107). d: posterior margin of sternite VIII strongly convex (Fig. 103); median lobe of aedeagus as in Figs. 104-105; apical lobe of paramere very long (Fig. 106). ?: posterior margin of sternite VIII rather weakly convex (Fig. 108); spermatheca as in Fig. 109. Comparative notes In external appearance (size, coloration, body shape, puncturation, microsculp- ture, antennal morphology, length of legs), the species somewhat resembles Cephalocousya nivicola (Thomson), but is distinguished by a more transverse pronotum, even more pronounced microsculpture of the forebody, longer elytra, and the completely different genitalia. From all other species of Cousya, it is sepa- rated by the matt appearance, the extremely fine puncturation of the forebody, as well as by the primary sexual characters. Distribution and bionomics The type locality is situated at the Ovitdagı Gecidi in the western parts of the Dogu Karadeniz Dagları (Fig. 71). The types were sifted from the roots of grass and herbs and from moss at an altitude of 2510 m. One of the specimens is teneral. Poromniusa crassa (Eppelsheim, 1883) Material examined: Izmir: 1 ex., 20km E Izmir, Nif Dagi, 38°23N, 27°22E, 1370-1400 m, sifted litter and grass roots under old Pinus, 26.XI1.2005, leg. AssınG (cAss). 48 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 This species is widespread, but rare, in northern, central, eastern, and southeast- ern Europe (AssING 1998). It is here recorded from Turkey for the first time. Ischnoglossa obscura Wunderle, 1990 Material examined: Aydin: 1 ex., Aydin, Dilek Dagi, S Kanyon, 37°40N, 27°11E, 670 m, 17.TV.2006, leg. BRACHAT & MEYBOHM (cAss). Ischnoglossa obscura had become known only from western and central Europe (WUNDERLE 1990). This is the first record from Turkey. Ocalea alutacea Eppelsheim, 1878 Material examined: Giresun: 1 ex. [det. SCHULKE], ca. 30km S Giresun, 40°36N, 38°27E, 1250 m, spruce forest with Rhododendron, sifted, 29.VI1.2006, leg. SCHULKE (cSch); 1 ex. [det. SCHULKE], ca. 30 km S Giresun, 40°35N, 38°27E, 1350 m, spruce forest with Rhododendron, 29.VII.2006, leg. SCHULKE (cSch). — Rize: 1 ex., 50km SSE Rize, W Sivrikaya, 40°41N, 40°39E, 2050m, natural Abies forest, litter and dead wood sifted, 1.VIII.2006, leg. AssınG (cAss); 1 ex., ca. 50km S Ardesen, Cat, 40°52N, 40°56E, 1240m, alder forest, sifted, 3.VIII.2006, leg. AsstNc (cAss). Ocalea alutacea had become known only from the West Caucasus (SMETANA 2004). The above specimens represent the first records from Turkey. Oxypoda (Oxypoda) acuminata (Stephens, 1832) Material examined: Ordu: 1 ex. [det. SCHULKE], Ordu, 25 km S Ordu, S Kabaduz, 40°49N, 37°54E, 990m, mixed forest with alder, spruce, bramble, ivy, 30.VII.2006, leg. SCHÜLKE (cSch). — Rize: 4 exs. [det. SCHULKE], 50 km SSE Rize, W Sivrikaya, 40°41N, 40°39E, 2050 m, natural fir forest, litter and dead wood sifted, 1.VIII.2006, leg. SCHULKE (cSch); 5 exs., ca. 50km S Ardesen, Cat, 40°52N, 40°56E, 1240 m, alder forest, sifted, 3. VIII.2006, leg. SCHULKE (cSch, cAss). In Turkey, this widespread Western Palaearctic species was previously known only from the European part (SMETANA 2004). Oxypoda (Oxypoda) subspectabilis n.sp. (Figs. 83, 110-118) Types Holotype 6: TR [22]-Ordu, 15 km $ Ordu, S Kabaduz, 990 m, mixed forest, 40°48'59N, 37°54'28E, 30.VII.2006, V. Assıng / Holotypus d Oxypoda subspectabilis sp. n. det. V. Ass- ING 2006 (cAss). Paratype @:same data as holotype (cAss). Etymology The name (Latin, adjective) refers to the fact that this species somewhat resembles a small specimen of O. spectabilis (Markel). Description Measurements (in mm) and ratios (holotype, paratype): AL: 1.92, 1.83; HL: 0.76, 0.77; HW: 0.74, 0.76; PW: 1.07, 1.07; PL: 0.86, 0.86; EL: 0.83, 0.80; EW: 1.24, 1.21; AW; 1.13, 1.13; TiL: 1.10, 1.04; TaL: 0.97, 0.97; ML: 0.83, -; TL: 6.3, 6.4; HL/HW: 1.02, 1.02; PW/HW: 1.45, 1.42; PW/PL: 1.25, 1.25; EL/PL: 0.96, 0.93; EW/PW: 1.15, 1.13; AW/EW: 0.91, 0.94; TiL/TaL: 1.14, 1.08. ASSING, STAPHYLINIDAE FROM TURKEY V 49 Figs. 110-118. Oxypoda subspectabilis n.sp. — 110. Habitus. 111. Forebody. 112. Antenna. 113. Head in lateral view. 114. Abdomen. 115-116. Median lobe of aedeagus in lateral and in ventral view. 117. Apical lobe of paramere. 118. Spermatheca. — Scale bars: 1 mm (110-111, 114), 0.5 mm (112-113), 0.2mm (115-118). Habitus as in Fig. 110. Coloration: head blackish brown; pronotum castaneous brown with narrowly reddish margins; elytra yellowish brown; abdomen dark brown, with the posterior margins of segments III-VI and the posterior 1/4-1/3 of segments VII-VIII yellowish; legs yellowish brown; antennae dark brown, with the basal 4 antennomeres reddish. Head approximately as wide as long; puncturation very shallow, ill-defined, and moderately dense, interstices on average approximately as wide as diameter of punc- tures; integument with distinct fine microreticulation; pubescence relatively short and depressed (Fig.111); eyes moderately large (Fig.113) and weakly projecting from lateral outline of head, slightly shorter than postocular region in dorsal view. Maxillary palpus with third joint slender, at least 3 times as long as wide. Antenna slender; antennomere III longer than II; IV oblong, distinctly shorter than III; V-X of subequal length and gradually increasing in width; X approximately as long as wide; XI approximately as long as the combined length of IX-X, without sexual di- morphism (Fig. 112). 50 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 Pronotum moderately transverse (see measurements and ratios PW/HW and PW/PL), maximal width in posterior half; posterior angles weakly marked, but no- ticeable (Fig. 111); puncturation, pubescence, and microsculpture similar to those of head; pronotal hypomera not distinctly visible in lateral view. Elytra distinctly wider and at suture slightly shorter than pronotum (see ratios EW/PW, EL/PL, and Fig.111); posterior margin sinuate near posterior angles; puncturation dense and fine, more distinct than that of pronotum; microsculpture shallower than that of pronotum; pubescence shorter and denser than that of prono- tum. Hind wings present and fully developed. Legs very slender (see measurements); metatarsus only slightly shorter than metatibia (see ratio TiL/TaL); metatarsomeres very long; metatarsomere I longer than the combined length of II-III, almost as long as the combined length of II-IV. Abdomen widest at segments IV/V, weakly tapering caudad (Fig. 114); punctura- tion and pubescence extremely fine and extremely dense; microsculpture shallow; posterior margin of tergite VII with palisade fringe; posterior margin of tergite VIII in both sexes strongly convex. 3: posterior margin of sternite VIII strongly convex and with very long marginal setae in the middle; median lobe of aedeagus of distinctive shape, rather massive, and with deeply incised ventral process (Figs. 115-116); apical lobe of paramere as in Fig. 117. Q: posterior margin of sternite VIII moderately convex and with shorter and stouter modified setae; spermatheca as in Fig. 118. Comparative notes Among other species of the subgenus, O. subspectabilis is characterised especially by the distinctive morphology of the median lobe of the aedeagus. In external ap- pearance (coloration, proportion) and the shape of the spermatheca, the new species is most similar to O. spectabilis, which has not been recorded from Turkey and from which it is additionally distinguished by smaller size (small specimens of O. spectabilis of similar size are very rare), a slightly less dense puncturation of the fore- body and the abdomen, shorter antennae (in O. spectabilis, antennomere X is ob- long), as well as by - both relatively and absolutely — shorter elytra. From other con- subgeners occurring in Turkey, O. subspectabilis is also readily separated by larger size, paler coloration, relatively shorter elytra, and the shape of the spermatheca. Distribution and bionomics The type locality is situated in the area to the south of Ordu, Ordu province (Fig. 83), at an altitude of approximately 1000 m. The specimens were sifted from the leaf litter of a mixed forest with alder, spruce, bramble, and ivy. Oxypoda (Thliboptera) micans Kraatz, 1855 Material examined: Aydin: 57 exs., ca. 20km NE Kuyucak, Bayrak Tepe, 37°58N, 28°33E, 850 m, N-slope, oak litter and grass sifted, 7.IV.2006, leg. Assıng, WUNDERLE (cAss, cWun); 22 exs., ca. 20km NE Kuyucak, Bayrak Tepe, 37°58N, 28°34E, 900 m, N-slope, oak litter and grass sifted, 7.1V.2006, leg. Assıng, WUNDERLE (cAss, cWun). This species was previously known only from Greece (AssING 2006c). It is here reported from Turkey for the first time. ASSING, STAPHYLINIDAE FROM TURKEY V Sl Oxypoda (Bessopora) ferruginea Erichson, 1839 Material examined: Ankara: 7 exs., SE Ankara, N-Elma Dagi, 1300 m, decaying and hollow Salix trunks, 31.X.1995, leg. Vrr (cAss). This rare species is widespread in the Western Palaearctic region, but was previ- ously unknown from Turkey (SMETANA 2004). Haploglossa villosula (Stephens, 1832) Material examined: Ankara: 2 exs., SE Ankara, N-Elma Dagi, 1300 m, hollow Salix trunks, 31.X.1995, leg. Vrr (cAss). This widespread and common Palaearctic species was only recently reported from Turkey (Antalya, Kahramanmaras) for the first time (AssınG 2006f). Dinusa smyrnensis n.sp. (Figs. 71, 119-122) Type Holotype @ [with two workers of Messor sp. attached to the same pin]: TR - Izmir [15], N Izmir, Yamanlar Dagi, pine forest, clearing, 940 m, 38°33'14N, 27°09'16E, 28.X11.2005, V. Assıng / Holotypus 2 Dinusa smyrnensis sp. n. det. V. AsstNG 2005 (cAss). Etymology The name (Latin, adjective) is derived from Smyrna, the ancient name of Izmir. Description Measurements (inmm) and ratios: AL: 1.43; HW: 0.79; PW: 1.28; PL: 0.69; EL: 0.62; EW: 1.36; TiL: 0.82; TaL: 0.60; TL: 4.8; PW/HW: 1.63; PW/PL: 1.85; EL/PL: 0.89; EW/PW: 1.06; TiL/TaL: 1.35. oe ' gent! id ry. 4 1 tree t. a oe Fe jahren er a! m mul Et an ‘aa ey, ee 119 121 122 Figs. 119-122. Dinusa smyrnensis n.sp. — 119. Habitus. 120. Forebody. 121. Abdomen. 122. Spermatheca. — Scale bars: 1 mm (119), 0.5 mm (120-121), 0.1 mm (122). 52 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 Habitus as in Fig. 119. Coloration: head blackish brown; pronotum dark brown with paler margins; elytra yellowish brown, near scutellum and anterior margin darkened; abdomen dark brown, with the apex (posterior 2/5 of segment VII and fol- lowing segments) yellowish brown; legs and antennae yellowish to reddish brown. Head transverse, with moderately dense fine puncturation and shallow mi- crosculpture; pubescence short and depressed, predominantly directed mediad; eyes not distinctly projecting from lateral outline of head in dorsal view, approximately as long as postocular region in dorsal view (Fig. 120). Antenna with antennomeres III-XI flattened; III distinctly oblong and much longer than II and IV; IV-X of subequal length and weakly oblong; XI slightly more than 1.5 times as long as wide, shorter than the combined length of IX. Pronotum strongly transverse (see measurements and ratios PW/HW and PW/PL), maximal width in posterior half; posterior angles rounded and ill-defined; puncturation and microsculpture similar to those of head; pubescence short and de- pressed, directed caudad along median line and predominantly diagonally latero- caudad in lateral areas (Fig. 120). Elytra at suture somewhat shorter than pronotum (see ratio EL/PL and Fig. 120); posterior margin strongly sinuate near posterior angles; puncturation slightly denser than that of head and pronotum; microsculpture slightly more distinct than that of head and pronotum; pubescence short, depressed, and directed caudad. Hind wings present and apparently fully developed. Metatarsus distinctly shorter than metatib- ia (see ratio TiL/TaL); metatarsomere I approximately as long as the combined length of H-IV. Abdomen widest at base, gradually decreasing in width posteriad (Fig. 121); ante- rior tergites without basal impressions; integument without microsculpture, inter- stices shiny; puncturation dense and fine, slightly sparser on posterior than on ante- rior tergites; posterior margin of tergite VII with distinct palisade fringe; posterior margin of tergite VIII strongly convex. 3: unknown. ?: spermatheca as in Fig. 122. Comparative notes The geographically closest congeners are Dinusa taygetana Eppelsheim from southern Greece and D. taurica Assing from central southern Anatolia. From the former, the new species is distinguished by a differently shaped pronotum (in D. taygetana with almost evenly rounded lateral margins, anteriorly less distinctly ta- pering), by much finer and sparser abdominal puncturation, and by the absence of microsculpture on the abdomen. From D. taurica, it is separated by a less transverse, anteriorly more strongly tapering, and more convex pronotum, slightly more dense- ly and more finely punctate pronotum and elytra, shorter metatarsi (in D. taurica approximately 0.7mm long, TiL/TaL: 1.25), and a more finely and sparsely punc- tured abdomen (particularly evident on tergite VII). For illustrations of the habitus and genitalia of D. taurica see AssiNG (2001a). Distribution and bionomics This myrmecophile is known from only one locality in the Yamanlar Dagi, to the north of Izmir, Izmir province (Fig. 71). The holotype was found in a nest of Messor sp., evidently its host, in a clearing of a pine forest at an altitude of 940 m. ASSING, STAPHYLINIDAE FROM TURKEY V 53 Megalogastria Bernhauer, 1901 (Figs. 123-135) Type species: Aleochara cingulata Eppelsheim, 1889. Redescription Body size moderately small. Head posteriorly with fine carina, not constricted; postgenae ventrally finely carinate. Antenna gradually incrassate apically, with dis- tinctly transverse preapical antennomeres and a weakly asymmetric antennomere XI (Fig. 125). Mouthparts distinctive; maxillary palpus with basal joint of reduced size, third joint rather large and approximately twice as long as wide, and fourth joint broad-based, of conical shape, pubescent, and apically constricted (Fig. 127); men- tum distinctly transverse, with two extremely long setae, with several additional se- tae of moderate length, and with two very short setae; labial palpus 3-jointed and conspicuously short, basal joint broad and about 1.5 times as long as wide, second joint very short and transverse, and apical joint short and stout; labium short and broad-based, apically incised and with several setae (Fig. 128); labrum distinctly transverse and weakly sclerotised; mandibles of the usual aleocharine morphology. Pronotum large, distinctly transverse and much wider than head; posterior angles completely rounded, obsolete (Fig. 124); pronotal hypomera not visible in lateral view; pubescence directed caudad along midline and more or less diagonally latero- caudad in lateral areas. Elytra in the type species of reduced length (Fig. 124), and hind wings reduced. Mesosternum anteriorly with median carina; mesosternal process apically acute and reaching approximately halfway between mesocoxae. Legs moderately slender; metatarsomere I elongated. Tarsal formula 5,5,5. Abdomen broad, widest at segments IV-V, segments VI-VIII distinctly tapering (Fig. 129); anterior impressions of segments III-V extremely shallow, almost obso- lete. 3: median lobe of aedeagus slender, of similar morphology as in some Oxypodi- ni and Aleocharini; internal sac with rather long flagellum and two apical sclerotised structures (Figs. 132-133); apical lobe of the paramere moderately elongated (Fig. 134). ?: morphology of spermatheca somewhat reminding of that of Aleochara species, with short and proximally truncate duct (Fig. 135). Discussion Megalogastria was originally described as a subgenus of Aleochara (Bernhauer, 1901). Assessing the phylogenetic affiliations of this remarkable taxon based on morphological characters alone is somewhat complicated. In various respects, it combines characters usually found in the Oxypodini with those confined to the Ale- ocharini. In body shape and general appearance (stout build, shape of pronotum) the genus is somewhat similar to species of Oxypoda Mannerheim, whereas the coloration, the complete absence of microsculpture of the whole body and the coarse puncturation of the abdomen give it some resemblance with some species of the subgenus Hete- rochara Mulsant & Rey of the genus Aleochara Gravenhorst. The mouthparts, which generally provide the most significant characters for the tribal assignment of Aleocharini and Oxypodini, pose another problem. The morphology of the labium and the maxilla, though in itself highly distinctive (and apparently highly derived) 54 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 does not clearly place the genus in either tribe. The fourth joint of the maxillary pal- pus is apically constricted, a condition usually not found in the Oxypodini and ap- proaching that of the Aleocharini. On the other hand, however, the apical part of the fourth joint does not represent a clearly separated pseudosegment as is generally the case in Aleocharini species. The shape (broad base, reduced length) and chaetotaxy (presence of several apical setae) of the ligula is to some extent similar to those found in some Aleochara, but the conspicuously short labial palpus lacks the pseudoseg- ment at the apex of the third joint. The morphology of the aedeagus does not present any conclusive evidence either way, although the general shape and the internal structures (long flagellum, shape of apical structures) are more similar to the condi- tion mostly found in Aleocharini. Finally, the morphology of the spermatheca dis- tinctly resembles that of Aleocharini and would be highly unusual for a member of Oxypodini. In conclusion, based especially on the morphology of the maxillary palpus, the shape and chaetotaxy of the ligula, the shape and internal structures of the aedeagus, as well as especially on the shape of the spermatheca, Megalogastria is assigned to the Aleocharini. However, in view of the conflicting evidence, this tribal placement can be considered only tentative. Other - e. g. molecular — characters are needed to ver- ify the correctness of this hypothesis. In any case, in view of the doubtful and isolat- ed phylogenetic position, Megalogastria is here attributed generic rank. Comparative notes Among other Aleocharini and also Oxypodini, Megalogastria is readily identified by the unique morphology of the mouthparts and the primary sexual characters. It is additionally characterised by the complete absence of microsculpture and the re- sulting shiny appearance, the coarse puncturation of the abdomen, the absence of distinct anterior impressions on the abdominal tergites III-V, as well as by the Oxy- poda-like body shape and the conspicuous coloration. Megalogastria cingulata (Eppelsheim, 1889) (Figs. 71, 123-135) Aleochara (Ceranota) cingulata Eppelsheim, 1889; ErPELSHEIM (1889: 164f.). Aleochara (Ceranota) Inteipennis Eppelsheim, 1889; EpPpELSHEIM (1889: 165f.). Type material examined: A. cingulata: Lectotype d, present designation: Smyrna, 25.11.86. Kors / 36 / Iuteipennis Epp. (immatur) / Aleochara cingulata Epp. / c. EpPELsH. STEIND. d. / Typus / Lectotypus d Aleochara cingulata Eppelsheim desig. V. AsstNG 2006 / Megalogastria cingulata (Eppelsheim) det. V. Assıng 2006 (NHMW). — Paralectotype 2: Smyrna, 25.III.86. Korg / 40 / cingulata Epp. Deutsch. ent. Zeit. 1889. p. 164 / c. ErpELsH. STEIND. d. / Typus / Paralectotypus Aleochara cingulata Eppelsheim desig. V. AssınG 2006 / Megalogastria cingulata (Eppelsheim) det. V. Assıng 2006 (NHMW). — A. luteipennis: Holotype 6: Turcia, MErkr. / 22 / Aleochara luteipennis Epp. / Inteipennis Epp. Deutsch. ent. Zeit. 1889. p. 165 / c. ErpELsH. STEIND. d. / Typus / Holotypus & Aleochara Inteipennis Ep- pelsheim rev. V. Assıng 2006 / Megalogastria cingulata (Eppelsheim) det. V. AssınG 2006 (NHMW). Additional material examined: Istanbul: 1 ex., “Belgrader Wald”, VIII.1974, leg. SCHUBERT (cAss). — Mugla: 2 exs., southern shore of Bafa lake, 37°29N, 27°29E, 24.111.2006, leg. MEY- BOHM (cAss); 2 exs., same locality, 19.-29.111.2006, leg. MEYBOHM (cAss). ASSING, STAPHYLINIDAE FROM TURKEY V 55 Comments The original description of A. cingulata is based on two syntypes from “Smyrna” (= Izmir), both of which EprELsHEIm (1889) erroneously believed to be females (“in 2 weiblichen Ex. aufgefunden”). Both type specimens are deposited in the Ep- PELSHEIM collection at the NHMW; the male is here designated as the lectotype. The original description of A. /uteipennis is explicitly based on a single specimen (“ein einziges ... Stück”), which consequently has holotype status, from an unspecified locality (“in der Türkei”). The synonymy of both names, which was established by BERNHAUER (1901), is here confirmed. Redescription (see also redescription of genus) Measurements (inmm) and ratios (range; n = 5): AL: 0.76-0.82; HL: 0.42-0.47; HW: 0.42-0.45; PW: 0.63-0.71; PL: 0.50-0.56; EL: 0.33-0.38; EW: 0.69-0.79; AW: 0.76-0.86; TiL: 0.51-0.57; TaL: 0.42-0.47; ML: 0.38-0.42; TL: 3.1-3.5; HL/HW: 1.00-1.03; PW/HW: 1.50-1.59; PW/PL: 1.27-1.39; EL/PL: 0.63-0.70; EW/PW: 1.04-1.11; AW/EW: 1.06-1.16; TiL/TaL: 1.19-1.27. Habitus as in Fig. 123. Usual coloration: distinctly bicoloured species, with the head and abdominal segments V-VII black, pronotum, elytra, and abdominal seg- ments III-IV bright reddish, and abdominal segment VIII reddish brown; legs and antennae reddish yellow. Whole body without microsculpture and very shiny. - The holotype of Aleochara luteipennis is somewhat darker, with the pronotum dark brown and with the anterior abdominal segments and the abdominal apex dark red- dish brown. Head approximately as wide as long; puncturation sparse and extremely fine; pu- bescence fine, moderately long, and depressed (Fig. 124); eyes moderately large (Fig. 126) and weakly projecting from lateral outline of head, slightly shorter than postocular region in dorsal view. Antenna relatively short and gradually incrassate apically; antennomeres I-III of subequal length; IV approximately as wide as long; V-X of increasing width and increasingly transverse; X approximately twice as wide as long; XI almost as long as the combined length of IX-X (Fig. 125). Pronotum distinctly transverse and wider than head (see measurements and ratios PW/PL and PW/HW); maximal width in posterior half, more strongly tapering cephalad than caudad; posterior angles rounded; puncturation fine, but more dis- tinct than that of head (Fig. 124). Elytra slightly wider and at suture shorter than pronotum (see measurements and ratios); posterior margin near posterior angles obliquely truncate; puncturation of similar density as that of pronotum, but more distinct and weakly granulose (Fig. 124). Hind wings reduced. Metatarsomere I conspicuously long, approximate- ly as long as the combined length of II-IV. Abdomen slightly wider than elytra; puncturation conspicuously coarse and moderately dense, slightly sparser on tergite VII than on anterior tergites (Fig. 129); posterior margin of tergite VII with narrow rudiment of a palisade fringe. 3: tergite VIII approximately as long as wide, its posterior margin weakly convex (Fig. 130); sternite VIII approximately as long as tergite VIII, its posterior margin obtusely angled in the middle (Fig. 131); median lobe of aedeagus with rather long flagellum and with pair of apical sclerotised structures in internal sac (Figs. 132-133); apical lobe of paramere as in Fig. 134. ?: posterior margin of tergite VIII convex; posterior margin of sternite VIII dis- 56 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 125 ali £81 Figs. 123-135. Megalogastria cingulata (Eppelsheim). - 123. Habitus. 124. Forebody. 125. An- tenna. 126. Head in lateral view. 127. Maxillary palpus. 128. Labium. 129. Abdomen. 130. Male tergite VIII. 131. Posterior part of male sternite VIII. 132-133. Median lobe of aedeagus in lateral and in ventral view. 134. Paramere. 135. Spermatheca. — Scale bars: 1 mm (123), 0.5mm (124, 126, 129), 0.2mm (125, 130-131), 0.1mm (127-128, 132-134), 0.05 mm (135). tinctly convex, almost obtusely angled in the middle, and with long marginal setae; spermatheca highly distinctive (Fig. 135). Comparative notes Megalogastria cingulata somewhat resembles some species of the subgenus Hete- rochara Mulsant & Rey of the genus Aleochara, ot Maurachelia Bernhauer, and even ASSING, STAPHYLINIDAE FROM TURKEY V 3% some species of the subgenus Mocyta Mulsant & Rey of Atheta Thomson, e.g. A. sanguinolenta (Wollaston). It is readily identified based on external characters alone, its oxypodoid body shape, the absence of microsculpture, the distinctively bi- coloured body, the pronounced puncturation of the abdomen, and the long metatar- somere I. For additional characters see the comparative notes below the redescrip- tion of the genus. Distribution and bionomics The species has become known only from Istanbul, Izmir, and Mugla provinces in western Turkey (BERNHAUER 1901, and material listed above) (Fig. 71). According to MEYBOHM (pers. comm.) the specimens from Mugla were sifted from Mastix lit- ter at little above sea-level. Pseudocalea messorphila n.sp. (Figs. 83, 136-146) Type Holotype 6 [with worker of Messor sp. attached to the same pin]: TR - Izmir [16], N Izmir, Yamanlar Dagi, 680 m, road margin, under stones, 38°32'42N, 27°09’42E, 28.X1I.2005, V. Assinc / Holotypus d Pseudocalea messorphila sp. n. det. V. AssınG 2005 (cAss). Etymology The name (adjective) alludes to the fact that the holotype was found in a nest of Messor sp. Description Measurements (inmm) and ratios: AL: 1.53; HL: 0.66; HW: 0.60; PW: 0.76; PL: 0.69; EL: 0.69; AW: 0.92; TiL: 0.89; TaL: 0.72; ML: 0.68; TL: 5.2; HL/HW: 1.10; PW/HW: 1.25; PW/PL: 1.09; EL/PL: 1.00; TiL/TaL: 1.23. Habitus as in Fig. 136. Coloration: head blackish brown; pronotum dark brown; elytra castaneous; abdomen blackish brown with the posterior margins of segments II-VI light brown; legs yellowish brown; antennae dark brown, with the basal 2 antennomeres light brown. Head oblong; puncturation rather shallow, moderately fine and moderately dense, interstices on average slightly wider than diameter of punctures; integument with distinct fine microreticulation; pubescence moderately long, pale, and suberect (Fig. 137); eyes moderately large (Fig. 139) and weakly projecting from lateral out- line of head, somewhat shorter than postocular region in dorsal view. Maxillary pal- pus with third joint distinctly dilated apically, approximately twice as long as wide, only slightly longer than the combined length of fourth joint and apical pseudoseg- ment. Antenna slender; antennomere III slightly longer than II; IV short and approximately as long as wide; V-X of gradually increasing size, weakly transverse; X only slightly wider than long; XI of ovoid shape and approximately as long as the combined length of IX-X (Fig. 138). Pronotum weakly transverse (see measurements and ratios PW/HW and PW/PL), maximal width in anterior half; posterior angles marked; puncturation very shallow and ill-defined, denser than that of head; microsculpture similar to that of head; pubescence long, pale, and depressed, directed cephalad along anterior !/; and caudad along posterior 2/3 of midline, and predominantly laterad in lateral areas (Fig. 137). 58 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 tom] 6 ee i 138 141 Figs. 136-146. Pseudocalea messorphila n.sp. — 136. Habitus. 137. Head and pronotum. 138. Antenna. 139. Head in lateral view. 140. Abdomen. 141. Male tergite VIII. 142-143. Me- dian lobe of aedeagus in lateral and in ventral view. 144-145. Apical part of median lobe in lat- eral and in ventral view. 146. Paramere. — Scale bars: 1 mm (136-137, 140), 0.5 mm (138-139), 0.2 mm (140-146). Elytra distinctly wider than and at suture approximately as long as pronotum (see ratio EL/PL and Fig. 136); posterior margin sinuate near posterior angles; punctura- tion more distinct than that of pronotum; microsculpture diagonally transverse, ASSING, STAPHYLINIDAE FROM TURKEY V 59 much shallower than that of head and pronotum; pubescence fine, predominantly directed caudad. Hind wings present and fully developed. Metatarsus somewhat shorter than metatibia (see ratio TiL/TaL); metatarsomere I longer than the com- bined length of I-III, but shorter than the combined length of II-IV. Abdomen with basal half subparallel, gradually tapering from segment V—VIII (Fig. 140); anterior impressions of tergites III-V distinct, that of tergite VI shallow; puncturation fine, distinct, and moderately dense; tergites without microsculpture, except for very indistinct traces on tergite VII; posterior margin of tergite VII with palisade fringe; posterior margin of tergite VIII distinctly concave (Fig. 141). 3: posterior margin of sternite VIII acutely pointed; median lobe of aedeagus slender and of distinctive shape (Figs. 142-145); paramere as in Fig. 146. ?: unknown. Comparative notes The genus previously comprised four species - one from the Eastern Palaearctic and three from the Western Palaearctic region - which may be distinguished using the following key: 1 Very large species (TL: approximately 7mm; PL: approx. 1.0mm; PW: approx. 1.1 mm), habitus as in Assinc (2006g: figs. 41-42); hind legs dark brown. Median lobe of aedeagus as illustrated by AssınG (2006g: figs. 49-51). — China: Yunnan. ......P. schuelkei Assing — Distinctly smaller species; all legs yellowish brown to rufous. Genitalia different. ...... 2 2 Smaller (TL approx. 3.7 mm) and paler species; pronotum, elytra, antennae, margins of ab- dominal terga, and abdominal apex ferrugineous, head and remainder of abdomen slightly darker; antennae shorter, with a transverse antennomere IV; elytra with distinctly granu- lose puncturation. Median lobe of aedeagus as in AssiNG (2002a: figs. 115-116). — Turkey (locality MOt-SpeClHed y= an ee olathe Teeth istt Potibas y ROM kates ar P. korbi (Bernhauer) — Mostly larger and darker species; antennomere IV at least as long as wide; elytral punc- turation not granulose. Genitaliaditterent: 29... Aue eee ako ete oe ole nen nen one elke 3 3 Pronotum brown to dark brown; pronotum distinctly transverse (PW/PL: 1.10-1.20) and with anterior angles less strongly bent ventrad, so that both anterior angles are visible simultaneously when viewed from straight above; elytra at suture usually slightly shorter than pronotum. Median lobe of aedeagus and spermatheca as in AssING & WUNDERLE (1998: figs. 10e-i). — Southern central, southern, and southeastern Europe. ............. gre ER 2.1. Sep. apt © pices oe RT Rt We cert A a, SREB a, ahs Ra el, P. brevicornis (Kraatz) — Pronotum usually blackish brown to black and less transverse (PW/PL: < 1.10), anterior angles strongly bent ventrad, so that they are not visible simultaneously when viewed from straight above; elytra at suture at least as long as pronotum. Genitalia different. ....... 4 4 Coloration darker, pronotum and elytra usually dark brown to blackish, base of antennae not distinctly paler than apical antennomeres, posterior margins of abdominal tergites only indistinctly paler; pubescence of head and pronotum sparse and suberect; antennae longer, antennomere IV at least weakly oblong; puncturation of abdomen very sparse. Median lobe of aedeagus and spermatheca as in AsstNG (2000: figs. 1a-b). - Southeastern Europe. sage Srila et Breed eh Ne P. angulata (Eppelsheim) — Coloration paler, pronotum and elytra brown, base of antennae distinctly paler than apical antennomeres, posterior margins of abdominal tergites distinctly paler; pubescence of head and pronotum depressed; antennae shorter, antennomere IV subquadrate (Fig. 138). Median lobe of aedeagus as in Figs. 142-145. — Western Anatolia (Fig. 83)...... P. messorphila n. sp. Distribution and bionomics The type locality is situated in the Yamanlar Dagi, to the north of Izmir, Izmir province (Fig. 83). The holotype was found in a nest of Messor sp. on a grassy road margin at an altitude of 680 m. Whether the species is really myrmecophilous or was 60 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 700 just accidentally close to the ant nest seems uncertain; morphological adaptations to an association with ants were not observed. Aleochara honesta Likovsky, 1973 Material examined: Gümüshane: 1 ex., ca. 50km SW Trabzon, E Zigana Geg., ca. 40°37N, 39°26E, 2500 m, 11.V1.1998, leg. SOLODOVNIKOV (cAss). — Rize: 1 ex., 40km SSW Hopa, source of Caglayan D., ca. 41°06N, 41°22E, 2700-2900 m, 25.V1.1998, leg. SOLODOVNIKOV (cAss); 1 ex., 30km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1800-1900 m, 26.V1.1998, leg. SOLODOVNIKOV (cAss). — Artvin: 2 exs., ca. 40km SW Artvin, source of Barhal river, 41°05N, 31°30E, 2400-2800 m, 23.-24.V1.1998, leg. SOLODOVNIKOV (cAss). SMETANA (2004) attributes A. honesta to RouBAL (1969), who first proposed this name as a replacement name for the preoccupied A. rufitarsis var. fortepunctata Roubal, 1911. According to Article 45.6.3 of the Code (ICZN 1999), however, A. honesta Roubal, 1969 is not an available name, since it was explicitly proposed as a variety (A. diversa v. honesta). It was LikovskyY (1973) who, in considering A. hones- ta a distinct species, made the name available. The species was previously known only from the West Caucasus (Likovsxy 1973) and is here recorded from Turkey for the first time. Aleochara ignipennis Fauvel, 1900 Material examined: Adana: 1 ex., NW Pozanti, Ceykavak Geg., 1600 m, 22.VII.1998, leg. BAYER (cAss). The above specimen represents the first record of this species from Turkey. Previ- ously, it had been reported from Armenia, Azerbaijan, and Iran (SMETANA 2004). Aleochara intricata Mannerheim, 1830 Material examined: Rize: 2 exs., 30km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1800-1900 m, 26.V1.1998, leg. SOLODOVNIKOV (cAss, cSol). — Artvin: 1 ex., ca. 40 km SW Artvin, source of Barhal river, 41°05N, 31°30E, 2400-2800m, 23.-24.V1.1998, leg. SOLODOVNIKOV (cAss). In Turkey, this species was previously only recorded from Van province (SCHEER- PELTZ 1958). Aleochara maculata Brisout de Barneville, 1863 Material examined: Gümüshane: 2 exs., ca. 50km SW Trabzon, 9-10km S Dikkaya, ca. 40°36N, 39°29E, 2000 m, 9.V1.1998, leg. SOLODOVNIKOV (cAss). — Erzurum: 3 exs., 35-40 km NW Tortum, Mescit Daglari, ca. 40°30N, 41°17E, 2600 m, 19.V1.1998, leg. SOLODOVNIKOV (cAss, cSol). The species is here reported from Turkey for the first time. Aleochara maculipennis (Baudi di Selve, 1857) Material examined: Konya: 1 ex., Beysehir, Camlik env., 29.1V.-1.V.2001, leg. Lona (cAss); 1 ex., Beysehir, Beysehir lake, Yesildag env., 5.-6.V.2001, leg. Lonay (cAss). The species was previously known only from Cyprus and “Syria” (SMETANA 2004). It is here reported from Turkey for the first time. ASSING, STAPHYLINIDAE FROM TURKEY V 61 Aleochara moesta Gravenhorst, 1902 Material examined: Erzurum: 15 exs., 30-45 km NNE Erzurum, Dumludagi, ca. 40°08N, 41°24E, 2200-2500 m, 14.V1.1998, leg. SOLODOVNIKOV (cAss, cSol). Aleochara moesta is widespread and common not only in the Palaearctic region, but has also been reported from the Oriental and the Ethiopian regions. It had been recorded from Turkish territory by SMETANA (2004), but without specification of lo- calities. Aleochara sparsa Heer, 1839 Material examined: Rize: 1 ex., 30 km SW Hopa, Caglayan river valley, ca. 41°09N, 41°22E, 1800-1900 m, 26.V1.1998, leg. SOLODOVNIKOV (cAss). The species is widespread and common in the Western Palaearctic region, but had been unknown from Turkey (SMETANA 2004). Aleochara stichai Likovsky, 1965 Material examined: Giresun: 1 ex., Giresun, ca. 40km S Giresun, N Kümbet, 40°34N, 38°26E, 1520 m, spruce forest with Rhododendron, 29.V11.2006, leg. AssınG (cAss). According to SMETANA (2004), the species was previously unknown from Turkey. Aleochara inconspicua Aubé, 1850 Material examined: Ankara: 8 exs., SE Ankara, N-Elma Dagi, 1300 m, hollow Salix trunk, 31.X.1995, leg. Vrr (cAss). The species was only recently reported from Turkey (Mugla, Kahramanmaras) for the first time (AssınG 2006f). 4 References AssING, V. (1997a): Review of the Palaearctic species of Autalia Leach in Samouelle, 1819 (Coleoptera, Staphylinidae, Aleocharinae). - Entomologische Blatter 93: 69-85. Assıng, V. (1997b): A revision of the Western Palaearctic species of Myrmecopora Saulcy, 1864, sensu lato and Eccoptoglossa Luze, 1904 (Coleoptera, Staphylinidae, Aleochari- nae, Falagriini). — Beiträge zur Entomologie, Berlin 47: 69-151. AssING, V. 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Author’s address: Dr. VOLKER Assıng, Gabelsbergerstr. 2, 30163 Hannover, Germany; e-mail: vassing.hann@t-online.de Manuscript received: 8.IX.2006, accepted: 6.X.2006. ISSN 0341-0145 Autoren-Richtlinien: http://www.naturkundemuseum-bw.de/stuttgart/schriften Schriftleitung: Dr. Hans-Peter Tschorsnig, Rosenstein 1, 70191 Stuttgart Gesamtherstellung: Gulde-Druck, 72072 Tübingen